Preface (G. E. Ball)
List of contributors
Chapter 1. Introduction (H. Turin, L. Penev & A.
Casale)
Why Carabus?
Some problems and
‘solutions’
Nomenclature in Checklist,
Key to the adults and Species accounts
Key to the larvae
General chapters
Subspecies
Mapping
Ecological transects
About Europe
Acknowledgements
SPECIAL PART
Chapter
2. Checklist (A. Casale & B. Březina)
Introduction
Checklist
Hybrids
Chapter
3. Key to the adults (A. Casale & O. L.
Kryzhanovskij)
Introduction
Key to the subgenera
01. Limnocarabus
02I . Morphocarabus + 02II. Trachycarabus
+ 04. Eucarabus
03. Carabus (s.str.)
05. Tachypus
06. Archicarabus
07. Hemicarabus
08. Diocarabus
09. Aulonocarabus
10. Mesocarabus
11. Orinocarabus
12. Oreocarabus
13. Tomocarabus
14. Eurycarabus
15. Pachystus
16. Ctenocarabus
17. Rhabdotocarabus
18. Hygrocarabus
19. Chaetocarabus
20. Platycarabus
21. Heterocarabus
22. Sphodristocarabus
23. Megodontus
24. Pachycranion
25. Iniopachys
26. Chrysocarabus (including 27. Chrysotribax)
28. Macrothorax
29. Lamprostus
30. Procrustes
31. Procerus
Chapter 4. Key to the larvae (E. Arndt & K.
Makarov)
Introduction
Larval morphology
Key to the second and third instar larvae
Key to the European subgenera
Key to selected subgenera
Chapter 5. Species accounts (H. Turin, L.Penev, A.
Casale, E. Arndt, Th. Assmann,
K. Makarov, D. Mossakowski, Gy. Szél & F. Weber)
Introduction
Abbreviations used
Explanations to the sections in the species
accounts
Species number and name
General literature: checklists and catalogues
Taxonomy
Geographical distribution, chorotypes and range
characteristics
Distribution in Europe
General works
Maps
Ecology and Biology
Conservation
Species accounts
01. Limnocarabus
02I . Morphocarabus + 02II. Trachycarabus
03. Carabus (s.str.)
04. Eucarabus
05. Tachypus
06. Archicarabus
07. Hemicarabus
08. Diocarabus
09. Aulonocarabus
10. Mesocarabus
11. Orinocarabus
12. Oreocarabus
13. Tomocarabus
14. Eurycarabus
15. Pachystus
16. Ctenocarabus
17. Rhabdotocarabus
18. Hygrocarabus
19. Chaetocarabus
20. Platycarabus
21. Heterocarabus
22. Sphodristocarabus
23. Megodontus
24. Pachycranion
25. Iniopachys
26. Chrysocarabus (including 27. Chrysotribax)
28. Macrothorax
29. Lamprostus
30. Procrustes
31. Procerus
GENERAL PART
Chapter 6. Biology and ecology (Th. Assmann)
Introduction
The life form of the genus Carabus
Reproduction and development
Reproductive systems
Sexual attraction and the power of aggregation
Copulation
Eggs
Larvae
Pupae and tenerals
Rearing
Activity
Daily rhythms
Annual rhythm
Locomotory activity and orientation
Abiotic factors
Climatic factors, temperature, humidity, and light
Chemical factors
Structural parameters ot the habitat
Biotic factors
Diet
Interspecific competition
Parasites and parasitoids
Predators and defence mechanisms
Population biology
Population ecology
Fluctuations of population size
Age structure
Power of dispersal
Differentiation and population history
Morphometric differentiation
Karyotypic differentiation
Differentiation at the level of single loci
Glacial fauna and postglacial history
Hybrids and hybrid zones
Chapter 7. Phylogeny (E. Arndt, M. Brückner, M.
Marciniak, D. Mossakowski & F. Prüser)
Introduction
Results
Analysis of morphological data
Analysis of molecular data
Discussion
Comparison of examined genital and larval
characters
Phylogenetic results
Monophyly of the genus Carabus Linné
Discussion of the monophyly of the subgroups
Phylogenetic relationships inferred from molecular
data
Conclusion – a tentative phylogenetic hypothesis
for the genus Carabus (s.l.)
Appendix
Chapter 8. Biogeography (L. Penev, A. Casale & H.
Turin)
Introduction
World distribution of Carabus
The general range and its limits
Patterns of diversity and specificity
Present-day distribution of Carabus in
Europe
Species composition, diversity and regional
distribution
Taxonomic structure
Chorology
Zoogeographical division of Europe based on the Carabus
fauna
Locality-based biogeography of Carabus
Outline of the ‘local fauna’ concept: an anastomosis
between
zoogeography and ecology
The dataset: local faunas of European Carabus
Variation in local species diversity
Types of species distribution across habitat and
space
Geographical variation in local species
compositions
Gradient analysis
Biogeographical insights into the phylogeny and
faunogenesis of Carabus
Classification, phylogeny, and historical
biogeography: an unfinished story
The scene: Europe and its evolution from the
Permian period to the present
The early Carabini in Europe
Origins of the Carabus species in Europe
The early history of Carabus in the light of
Palaearctic-Nearctic connections
Biogeographical patterns and underlying processes
Geographical history of European Carabus species:
a scenario
Appendix 8.1. Distribution, taxonomic and
chorological characteristics of
Carabus species in
Europe
Appendix 8.2. Distribution map of 100 local faunas
and schematic pictures
of all local gradients (=transects)
Transects T-01-100
Chapter 9. Conservation biology (Th. Assmann)
Decline – examples and outline of the problem
What shall we protect? A population-biological
approach
Reasons for the decline of carabid populations
Habitat destruction
Fragmentation
Insecticides
Small population size
Collecting
Stopping the loss – preservation and sustainable
conservation
MVP, genetic monitoring
Habitat corridors
Reintroduction projects
Programmes of crop edges and field enhancement
Action plans for conservation
Conclusions
Chapter 10. Carabus evaluations (H. Turin, L. Penev, A.
Casale & E. Arndt)
Introduction
Chapter 2 – Checklist
Chapter 3 – Key to the adults
Chapter 4 – Key to the larvae
Chapter 5 – Species accounts
Chapter 6 – Biology and ecology
Chapter 7 – Phylogeny
Chapter 8 – Biogeography
Chapter 9 – Conservation
Conclusions
Glossary
Addendum
References
Author’s index
Taxonomic index