Latest Articles from Deutsche Entomologische Zeitschrift Latest 100 Articles from Deutsche Entomologische Zeitschrift https://dez.pensoft.net/ Tue, 19 Mar 2024 06:56:16 +0200 Pensoft FeedCreator https://dez.pensoft.net/i/logo.jpg Latest Articles from Deutsche Entomologische Zeitschrift https://dez.pensoft.net/ New genus and species of lice in the Oxylipeurus-complex (Phthiraptera, Ischnocera, Philopteridae), with an overview of the distribution of ischnoceran chewing lice on galliform hosts https://dez.pensoft.net/article/111874/ Deutsche Entomologische Zeitschrift 71(1): 85-109

DOI: 10.3897/dez.71.111874

Authors: Daniel R. Gustafsson, Chunpo Tian, Mengjiao Ren, Zhu Li, Xiuling Sun, Fasheng Zou

Abstract: Here, we describe a new genus of lice (Phthiraptera, Ischnocera) in the Oxylipeurus-complex, parasitising galliform hosts in the genera Tragopan Cuvier, 1829. This genus, Pelecolipeurus gen. nov., is separated from other members of the complex by the unique shape of the male subgenital plate and stylus, the male genitalia and other characters. The only previously-known species in the genus is Lipeurus longus Piaget, 1880, which is here tentatively re-described as Pelecolipeurus longus (Piaget, 1880), based on specimens from a non-type host, Tragopan temminckii (Gray, 1831). In addition, we describe a new species, Pelecolipeurus fujianensis sp. nov., based on specimens from Tragopan caboti (Gould, 1857). An overview of the distribution patterns of ischnoceran lice on galliforms is presented, which suggests that host phylogeny, host biogeography and host biotope, as well as elevation of host range, may all be important factors that have structured louse communities on landfowl. We transfer the genus Afrilipeurus from the Oxylipeurus-complex to the Lipeurus-complex and include an emended key to the Oxylipeurus-complex.

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Research Article Tue, 12 Mar 2024 09:36:08 +0200
Integrated taxonomy, biology and biogeography of the Afrotropical genus Xyloctonus (Coleoptera, Curculionidae, Scolytinae) https://dez.pensoft.net/article/116185/ Deutsche Entomologische Zeitschrift 71(1): 67-84

DOI: 10.3897/dez.71.116185

Authors: Bjarte H. Jordal

Abstract: The peculiar Afrotropical bark beetle genus Xyloctonus Eichhoff, 1872 is revised and its biology described. Several unusual morphological features reflect adaptations to predator avoidance as they are highly exposed during mating externally on tree trunks and branches. Observations invariably indicate that males and females abandon the nest under bark at an early stage of progeny, the males already before eggs hatch, potentially engaging in subsequent additional matings. Most species have a clear preference for host plants in the plant family Sapotaceae. Although the genus is broadly distributed in forested parts of Africa, Madagascar and Mauritius, most species are found in the eastern part of this range. A Bayesian biogeographical analysis revealed a possible origin of the genus in Madagascar in the early Eocene, with subsequent colonisation of the southern African region in late Eocene. This contrasts with the closely-related xyloctonine genus Ctonoxylon Hagedorn, 1910, which is of western Congolian ancestry and more recently reached Madagascar multiple times during late Miocene. Two new species are described: Xyloctonus magnus sp. nov. from Madagascar and X. genieri sp. nov. from Burkina Faso. Synonyms are proposed for X. subcostatus Eggers, 1939 (= X. striatus Eggers, 1939) and X. scolytoides Eichhoff, 1872 (= X. latus Eggers, 1922). Identification to species is provided in a key illustrated with photographs of most species.

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Research Article Fri, 1 Mar 2024 14:53:57 +0200
Two new species of the genus Agramma (Hemiptera, Heteroptera, Tingidae) from small islands of Japan, with an illustrated key to the Japanese species of the genus https://dez.pensoft.net/article/108270/ Deutsche Entomologische Zeitschrift 71(1): 49-65

DOI: 10.3897/dez.71.108270

Authors: Jun Souma

Abstract: The present study describes two new species of the monocotyledon-feeding lace bugs of the genus Agramma Stephens, 1829 (Hemiptera, Heteroptera, Tingidae, Tinginae, Tingini) from small islands of Japan. The first is A. (A.) izuense sp. nov., which was recorded as A. (A.) japonicum (Drake, 1948) from Hachijo Island, the Izu Islands, in a previous study, and is considered an independent species here based on morphological characteristics and molecular data. The second is A. (A.) keramense sp. nov., which has a remarkable spineless head and was discovered from Aka and Geruma islands, Kerama Group, the Ryukyu Islands. Consequently, the following four species of Agramma were recognized in Japan: A. (A.) abruptifrons Golub, 1990, A. (A.) izuense sp. nov., A. (A.) japonicum, and A. (A.) keramense sp. nov. Only dozens of submacropterous morphs were confirmed in these two species in the present study, suggesting that both new species are flightless. In addition, an illustrated key for the identification of the four species from Japan and the host plant relationships of the two new species are provided.

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Research Article Tue, 30 Jan 2024 12:18:57 +0200
Forest leaf litter beetles of Taiwan: first DNA barcodes and first insight into the fauna https://dez.pensoft.net/article/112278/ Deutsche Entomologische Zeitschrift 71(1): 17-47

DOI: 10.3897/dez.71.112278

Authors: Fang-Shuo Hu, Emmanuel Arriaga-Varela, Gabriel Biffi, Ladislav Bocák, Petr Bulirsch, Albert František Damaška, Johannes Frisch, Jiří Hájek, Peter Hlaváč, Bin-Hong Ho, Yu-Hsiang Ho, Yun Hsiao, Josef Jelínek, Jan Klimaszewski, Robin Kundrata, Ivan Löbl, György Makranczy, Keita Matsumoto, Guan-Jie Phang, Enrico Ruzzier, Michael Schülke, Zdeněk Švec, Dmitry Telnov, Wei-Zhe Tseng, Lan-Wei Yeh, My-Hanh Le, Martin Fikáček

Abstract: We report the publication of 953 DNA barcodes of forest leaf litter beetles from central Taiwan, in total representing 334 species of 36 beetle families. This is the first bulk of data from the Taiwanese Leaf Litter beetles project focused on uncovering the under-explored diversity of leaf litter beetles across Taiwan. Based on these data, we provide the first records of the following taxa for Taiwan: family Sphindidae (genus Aspidiphorus Ziegler, 1821); tribes Trichonychini, Ctenistini, and Bythinoplectini (all Staphylinidae: Pselaphinae); genera Gyrelon Hinton, 1942, Thyroderus Sharp, 1885, Cautomus Sharp, 1885 (all Cerylonidae), Dermatohomoeus Hlisnikovský, 1963 (Leiodidae), Paraploderus Herman, 1970 (Staphylinidae: Oxytelinae), Thinocharis Kraatz, 1859 (Staphylinidae: Paederinae), Cephennodes Reitter, 1884, Napoconnus Franz, 1957 (both Staphylinidae: Scydmaeninae), Bicava Belon, 1884 (Latridiidae), Otibazo Morimoto, 1961, Seleuca Pascoe, 1871 and Acallinus Morimoto, 1962 (all Curculionidae); species Oodes (Lachnocrepis) japonicus (Bates, 1873) (Carabidae: Licininae), Drusilla obliqua (Bernhauer, 1916) (Staphylinidae: Aleocharinae) and Coccotrypes advena Blandford, 1894 (Curculionidae: Scolytinae). The records of Anapleus Horn, 1873 (Histeridae) and Batraxis Reitter, 1882 (Staphylinidae: Pselaphinae) have been confirmed. The male of Sivacrypticus taiwanicus Kaszab, 1964 (Archeocrypticidae) is described for the first time. Gyrelon jenpani Hu, Fikáček & Matsumoto, sp. nov. (Cerylonidae) is described, illustrated, and compared with related species. DNA barcodes associated larvae of 42 species with adults, we are concisely illustrating some of these: Oodes japonicus, Perigona cf. nigriceps Dejean, 1831 (both Carabidae), Ptilodactyla sp. (Ptilodactylidae), Maltypus ryukyuanus Wittmer, 1970 (Cantharidae), Drusilla obliqua, Myrmecocephalus brevisulcus (Pace, 2008), Diochus sp., Mimopinophilus sp. (all Staphylinidae), Stelidota multiguttata Reitter, 1877, Lasiodites inaequalis (Grouvelle, 1914) (both Nitidulidae), Lagria scutellaris Pic, 1910, and Anaedus spinicornis Kaszab, 1973 (both Tenebrionidae). We also report the first cases of Rickettsia infections in Scydmaeninae and Pselaphinae. All data (sequences, metadata, and voucher photos) are made public in BOLD database and in a Zenodo Archive.

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Research Article Mon, 8 Jan 2024 15:53:39 +0200
Two new genera and four new species of Neotropical Eccritotarsini (Heteroptera, Miridae, Bryocorinae) https://dez.pensoft.net/article/104130/ Deutsche Entomologische Zeitschrift 71(1): 1-15

DOI: 10.3897/dez.71.104130

Authors: Eugenia Minghetti, Sara Itzel Montemayor, Pablo Matías Dellapé

Abstract: In this study two new genera and four new species of Eccritotarsini (Hemiptera: Heteroptera: Miridae: Bryocorinae) are described. Thomasomiris gen. nov. is established to accommodate Thomasomiris setosus sp. nov. (from Panama) and Egerocoris gen. nov., is described for E. ecuatorianus sp. nov. (the type species), E. dimorphus sp. nov. (both from Ecuador) and E. chaparensis sp. nov. (from Bolivia). Adult habitus and male genitalia photographs are provided for each species. A key to species is also provided and the affinities of the new genera are discussed.

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Research Article Mon, 8 Jan 2024 15:52:37 +0200
Two new species of Deuterophlebia Edwards, 1922 from Southwestern China (Diptera, Deuterophlebiidae) https://dez.pensoft.net/article/108096/ Deutsche Entomologische Zeitschrift 70(2): 387-401

DOI: 10.3897/dez.70.108096

Authors: Xuhongyi Zheng, Changfa Zhou

Abstract: The family Deuterophlebiidae is considered the most plesiomorphic Diptera family, with striking morphology and life history. In this study, we provide detailed descriptions and figures of two new species from Southwestern China: Deuterophlebia pseudopoda sp. nov. and Deuterophlebia pachychaeta sp. nov., along with genetic distances of COI sequences between the seven known Chinese species of Deuterophlebia Edwards, 1922. The two new species can be identified by the adults’ male terminalia, head structures, male and female antennae and mesothoracic spines or abdominal projections of the pupae. As a result, China is currently the country with the highest Deuterophlebia species diversity.

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Short Communication Mon, 23 Oct 2023 10:06:25 +0300
Taxonomic study of the tribe Campsomerini (Hymenoptera, Scoliidae) from northern Vietnam, with the description of a new species and a checklist of Vietnamese scoliid wasps https://dez.pensoft.net/article/101152/ Deutsche Entomologische Zeitschrift 70(2): 369-385

DOI: 10.3897/dez.70.101152

Authors: Phong Huy Pham, Cornelis van Achterberg

Abstract: The tribe Campsomerini (Hymenoptera: Scoliidae) from northern Vietnam was studied, resulting in the finding of three species and six subspecies belonging to five genera. Three genera (Megacampsomeris Betrem, 1928 (with M. shillongensis (Betrem, 1928)), Micromerialla Betrem, 1964 (with M. marginella marginella (KIug, 1810)) and Phalerimeris Betrem, 1967 (with P. phalerata phalerata (de Saussure, 1858))) and three species (Sericocampsomeris flavomaculata Gupta & Jonathan, 1989, Campsomeriella (Annulimeris) annulata (Fabricius, 1793) and C. (Campsomeriella) collaris (Fabricius, 1775)) are recorded for the first time from Vietnam. One new species, Sericocampsomeris vietnamica Pham & van Achterberg, sp. nov. is described and illustrated. Identification keys to the genera, species and subspecies of the tribe Campsomerini from northern Vietnam and to the species of Sericocampsomeris Betrem, 1941 are presented. Lastly, we provide a checklist with 13 species and subspecies of Scoliidae from Vietnam.

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Research Article Fri, 6 Oct 2023 10:17:49 +0300
Comparative morphology of the larval mouthparts among six species of Notodontidae (Insecta, Lepidoptera), with discussions on their feeding habits and pupation sites https://dez.pensoft.net/article/107431/ Deutsche Entomologische Zeitschrift 70(2): 357-368

DOI: 10.3897/dez.70.107431

Authors: Jia-Xin Liu, Lu Jiang

Abstract: Larval mouthparts are significant organs for the individual development, morphologically related with feeding habits, and providing valuable characters for taxonomy and phylogenetic analysis. In previous studies, larval mouthparts revealed two identifying characters of Notodontidae. However, the evolutionary driving force and exact definition of these structures remain unsatisfactory. In this study, the larval mouthparts of Euhampsonia cristata (Butler, 1877), Fentonia ocypete (Bremer, 1861), Phalera assimilis (Bremer & Grey, 1853), Nerice davidi Oberthür, 1881, Cerura erminea (Esper, 1783) and Furcula furcula (Clerck, 1759) are morphologically observed and compared using scanning electron microscopy (SEM). The larval mouthparts of the six species are commonly equipped with paired maxillary sacs, congruent with the previous descriptions. However, the larval mouthparts of N. davidi are peculiar for bearing toothed mandibles, providing an exception of Notodontidae. Otherwise, the mouthparts exhibit morphological differences on mandibles, spinnerets, labral notches, and setal arrangements among the six species. The morphological diversity and the related feeding and pupation habits are briefly discussed.

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Research Article Fri, 29 Sep 2023 15:36:56 +0300
Phylogeny of genus Sichuana Shen & Yin, 2020 (Orthoptera, Tettigoniidae, Tettigoniinae) with four new species from Sichuan, China https://dez.pensoft.net/article/105870/ Deutsche Entomologische Zeitschrift 70(2): 337-355

DOI: 10.3897/dez.70.105870

Authors: Jun-Jie Gu, Chengjie Zheng, Su-Rong Jiang, Yanli Yue

Abstract: Four new species of Sichuana Shen & Yin, 2020 are described based on morphological comparison and molecular analysis: S. planicercata sp. nov., S. curvicercata sp. nov., S. longilamina sp. nov. and S. magnicerca sp. nov. Specimens showed some intraspecific variation of male tegmina and subgenital plates. The genes COI and 16S were used to analyze the genetic distance between species and COI was used to analyze the phylogenetic relationship of Sichuana.

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Research Article Fri, 29 Sep 2023 11:28:58 +0300
Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini) https://dez.pensoft.net/article/107425/ Deutsche Entomologische Zeitschrift 70(2): 311-335

DOI: 10.3897/dez.70.107425

Authors: Arnaud Faille, Sylvia Hofmann, Yeshitla Merene, David Hauth, Lars Opgenoorth, Yitbarek Woldehawariat, Joachim Schmidt

Abstract: The trechine beetle fauna (Coleoptera, Carabidae) of the Ethiopian Highlands is known to be highly diverse in species, and many species groups were recognized to be characterized by unusual character states of external and genital morphology. Earlier authors described several genera and subgenera of Ethiopian Trechina endemic to certain high mountains of the country. However, the relationships of these species groups and their evolutionary history are unknown so far. Here, we present the first molecular phylogenetic analysis of Ethiopian Trechina, detect several synonymic names under Trechus sensu lato, and introduce two new species groups to the country’s fauna: the monotypic genus Baehria Schmidt & Faille, gen. nov., with the type species B. separata sp. nov. from Mt. Choke in northern Ethiopia, and the Trechus subgenus Abunetrechus Schmidt & Faille, subgen. nov., with the type species T. bipartitus Raffray, 1885; this subgenus includes three species of northern Ethiopia. We show that the composition of the Ethiopian fauna is based on multiple events of immigration, which started simultaneously with or some million years after the Oligocene-Early Miocene orogenic events north and south of the Rift Valley. Our results support the habitat island hypothesis for the evolution of the Ethiopian highland fauna. We found no evidence for an alternative hypothesis assuming a close connection of the Trechina immigration to Ethiopia and Pleistocene cooling. We, thus, conclude that the geomorphological development rather than the climatic changes are the main drivers of the diversification of the high-altitude Trechina fauna in Ethiopia.

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Research Article Wed, 27 Sep 2023 17:44:12 +0300
A morphology-based revision and phylogenetic analysis of the Pterostichus macrogenys species group (Coleoptera, Carabidae) and implications for differentiation of the species group https://dez.pensoft.net/article/107322/ Deutsche Entomologische Zeitschrift 70(2): 291-310

DOI: 10.3897/dez.70.107322

Authors: Kôji Sasakawa, Yoshiji Mitsuduka

Abstract: The Pterostichus macrogenys species group is an endemic subterranean Japanese carabid clade that provides intriguing material for studying morphological differentiation, speciation and interspecific relationships. However, its diversity remains not fully explored. We investigated specimens from northern Tohoku District, an area where knowledge of this species group is notably limited. Our research led to the description of three new species: P. namahage sp. nov., P. kamurosanus sp. nov. and P. atsumidakensis sp. nov. We also updated distribution records for three known species: P. asahinus Habu & Baba, 1960; P. kitakamisanus Sasakawa, 2005; and P. chokaisanus Sasakawa, 2009. This report includes both the expanded distribution ranges of the known species and the discovery of a new sympatric species pair (P. chokaisanus and P. asahinus). We conducted a morphological phylogenetic analysis of all but one species, for which no male specimens were available, accounting for a total of 42 species within the group. The resulting phylogenetic tree implies that the initial differentiation of this species group originated on the Sea of Japan side, in the northern part of their current distribution, followed by dispersion to other areas and subsequent differentiation. Additionally, our findings indicate that sympatric species of varying body sizes are distantly related phylogenetically. These insights into the differentiation process align with regional distribution patterns of species-level diversity and sympatric sites.

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Research Article Thu, 17 Aug 2023 18:01:47 +0300
Taxonomic review of the genus Sinopanorpa Cai & Hua, 2008 (Mecoptera, Panorpidae) with descriptions of two new species https://dez.pensoft.net/article/104621/ Deutsche Entomologische Zeitschrift 70(2): 283-290

DOI: 10.3897/dez.70.104621

Authors: Yuan Hua, Kai Gao, Lianxi Xing

Abstract: The scorpionfly genus Sinopanorpa Cai & Hua, 2008 is endemic to the mountain regions of central China, currently consisting of four species: Sinopanorpa tincta (Navás, 1931), S. digitiformis Huang & Hua, 2008, S. nangongshana Cai & Hua, 2008, and S. baokangensis Wang, 2021. Here, the genus is taxonomically reviewed, with descriptions of two new species: Sinopanorpa minshanicola sp. nov. from the Minshan Mountains and Sinopanorpa shennongjiaica sp. nov. from the Shennongjia Mountains, increasing the species number of the genus to six. An updated key to species of Sinopanorpa is provided.

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Research Article Thu, 27 Jul 2023 16:20:36 +0300
An “American” silkmoth endemic to Himalayas, part I: life history and natural distribution of Antheraea compta Rothschild, 1899 (Lepidoptera, Saturniidae) https://dez.pensoft.net/article/102952/ Deutsche Entomologische Zeitschrift 70(2): 261-282

DOI: 10.3897/dez.70.102952

Authors: Zhengyang Liu

Abstract: Females of the uncommon Antheraea compta Rothschild, 1899 were collected at the natural habitat in the Tibetan Sub-Himalayas, descendants of which were reared on Quercus yunnanensis (Fagaceae) successfully, with all the preimaginal instars recorded morphologically. Correlated characters revealed a close relationship between this taxon and New World Antheraea spp., suggesting more attention and protection towards this key species is necessary in the future.

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Research Article Mon, 24 Jul 2023 13:01:56 +0300
Bush-crickets with very special ears and songs – review of the East African Phaneropterinae genus Dioncomena Brunner von Wattenwyl, 1878, with notes on its biogeography and the description of new species https://dez.pensoft.net/article/100804/ Deutsche Entomologische Zeitschrift 70(2): 221-259

DOI: 10.3897/dez.70.100804

Authors: Claudia Hemp, Fernando Montealegre-Z, Charlie Woodrow, Klaus-Gerhard Heller

Abstract: This study focuses on the genus Dioncomena and its acoustics, particularly the unique songs produced by male Dioncomena that consist of several distinct elements in a fixed sequence, culminating in a coda that typically elicits a response from a receptive female. We also examine the inflated pronotal lobes, which we term prebullae, that are prominently developed in some Dioncomena species but not in others. We discuss the role of prebullae in the context of acoustic communication in Dioncomena and other related Phaneropterini genera that have similar lateral pronotal lobes. We found that prebullae size is correlated with habitat distribution, with larger prebullae occurring in isolated species while aggregation-prone species have smaller or less pronounced prebullae. Using micro-computer tomography we show sexual dimorphism in the 3D geometry of the acoustic tracheae, being larger in the male. Interestingly, the tracheae are coupled by a septum, like in field crickets, which suggests potential cross talk. We define three groups of Dioncomena based on altitude preferences, ecology, color patterns, and songs: the jagoi-, tanneri-, and ornata-groups. We describe the songs of several species, including newly identified species such as D. flavoviridis sp. nov., D. magombera sp. nov., D. ngurumontana sp. nov., D. sanje sp. nov., D. tanneri, D. versicolor sp. nov., and D. zernyi. We also provide information on the nymphs, development time, and mating behavior of various species reared in the laboratory, shedding light on their phenology and adaptations to their habitats.

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Research Article Mon, 10 Jul 2023 12:22:51 +0300
Change of Editor-in-Chief https://dez.pensoft.net/article/107541/ Deutsche Entomologische Zeitschrift 70(2): 219-219

DOI: 10.3897/dez.70.107541

Authors: Matthias Seidel, Dominique Zimmermann

Abstract:

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Editorial Fri, 23 Jun 2023 10:04:08 +0300
The snakeflies of the Mediterranean islands: review and biogeographical analysis (Neuropterida, Raphidioptera) https://dez.pensoft.net/article/101559/ Deutsche Entomologische Zeitschrift 70(1): 175-218

DOI: 10.3897/dez.70.101559

Authors: Horst Aspöck, Ulrike Aspöck

Abstract: The Mediterranean region representsone of the hotspots of biodiversity of many organisms and among these also of Raphidioptera. With about 250 known species world-wide Raphidioptera is the smallest order of extant holometabolous insects. The Mediterranean region harbors 113 species (= >45% of the world fauna), and of these 33 (= >13% of the world fauna) have been found on islands of the Mediterranean Sea. In the course of the past 50 years most of the larger islands have been intensively explored for Raphidioptera, particularly all islands (except the Baleares) which had no connection to the mainland throughout the whole Pleistocene. Altogether, 11 species of Raphidioptera (9 Raphidiidae, 2 Inocelliidae) are endemic to (usually)one, rarely to a few Mediterranean islands: Phaeostigma (Ph.) euboica (Euboea), Ph. (Aegeoraphidia) prophetica (Rhodes), Ph. (Ae.) karpathana (Karpathos), Ph. (Ae.) biroi (Crete), Ph. (Superboraphidia) minois (Crete), Subilla principiae (Sardinia), S. colossea (Rhodes), Raphidia (R.) peterressli (Chios), R. (R.) ariadne (Crete), Fibla (F.) maclachlani (Corsica, Sardinia, Sicily), F. (Reisserella) pasiphae (Crete). In the Aegean Sea there is a remarkable difference between the eastern and the western islands marked by the so-called Rechinger line based on results of the analysis of the flora. The Raphidioptera confirm this line impressively. It is rather unlikely that further Raphidioptera species endemic to Mediterranean islands are still to be detected – with one exception: the Baleares. A few additional species known from various parts of the continents surrounding the Mediterranean Sea will probably be found on islands so that a total number may be around 40, possibly around 45 species. The possible ways and times of colonization of the islands by Raphidioptera are discussed. The present paper may serve as a basis for studies on natural and particularly on anthropogenic dispersals of snakeflies from island to island, from mainland to islands, and from islands to the mainland in connection with phylogeographic investigations.

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Research Article Wed, 3 May 2023 15:07:38 +0300
Resolving inconsistencies between Plötz’s descriptions and presumed type specimens of some Hesperiidae (Lepidoptera) https://dez.pensoft.net/article/98280/ Deutsche Entomologische Zeitschrift 70(1): 159-174

DOI: 10.3897/dez.70.98280

Authors: Jing Zhang, Qian Cong, Leina Song, Jinhui Shen, Théo Léger, Gerardo Lamas, Olaf H. H. Mielke, Nick V. Grishin

Abstract: Comparing specimens curated in MfN as primary types of Hesperiidae names proposed by Carl Plötz with the original descriptions and unpublished drawings reveals a number of inconsistencies that we address. Lectotypes are designated for Telegonus labriaris Butler, 1877, Eudamus jalapus Plötz, 1881, and Apaustus interpunctata Plötz, 1884. Neotypes are designated for Netrocoryne seneca Plötz, 1882 and Hesperia irma Plötz, 1882. Hesperia ulphila Plötz, 1883 is treated as a nomen dubium. As a result of these designations, the following are junior objective synonyms: Netrocoryne seneca Plötz, 1882 of Telegonus labriaris Butler, 1877 and Hesperia irma Plötz, 1882 of Pamphila irma Möschler, 1879, the latter two names being homonyms. In all these instances, the original descriptions are satisfied, and, except for A. interpunctata, the current application of these names is unaffected. Taxonomically, Callimormus diaeses Schaus, 1902, stat. rest. is a valid species, not a synonym of A. interpunctata, which is a junior subjective synonym of Eutocus vetulus (Mabille, 1883).

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Research Article Mon, 3 Apr 2023 15:39:55 +0300
The genus Anacaena Thomson from the Ryukyu Archipelago of Japan (Coleoptera, Hydrophilidae) https://dez.pensoft.net/article/96994/ Deutsche Entomologische Zeitschrift 70(1): 143-157

DOI: 10.3897/dez.70.96994

Authors: Yûsuke N. Minoshima, Yuuki Kamite, Martin Fikáček

Abstract: We review the genus Anacaena Thomson, 1859 from the Ryukyu Archipelago, southern Japan. Three aquatic species are recognised: A. torikaii sp. nov. from Amami-ôshima Island, A. okinawana sp. nov. from Okinawa-jima Island and Kerama Islands, and A. kumejimana sp. nov. from Kumejima Island. All three species are very similar, with the morphology of the aedeagus being essential for a reliable identification. Dorsal colouration is also useful as a diagnostic character, despite some variation within species. We observe a possible geography-based variation between A. okinawana from Okinawa-jima I. and the neighbouring Kerama Is., but we treat both populations as conspecific based on genital morphology. Anacaena kumejimana and A. okinawana share many morphological characters possibly indicating their close relationship. We compare the endemism of aquatic Hydrophilidae in the Ryukyu Archipelago to that in other groups of aquatic beetles: the proportion of endemic species is higher in aquatic Hydrophilidae than in Dytiscidae, but much lower than in stream-inhabiting Hydraenidae and Elmidae. A list of Japanese species of Anacaena and a key to the Japanese species of the genus are provided.

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Research Article Fri, 24 Mar 2023 10:45:37 +0200
Caribbean pygmy jumping leaves (Tetrigidae, Cladonotinae, Choriphyllini) https://dez.pensoft.net/article/98982/ Deutsche Entomologische Zeitschrift 70(1): 129-141

DOI: 10.3897/dez.70.98982

Authors: Josip Skejo, Sheyla Yong, Domagoj Bogić, Niko Kasalo

Abstract: The tribe Choriphyllini Cadena-Castañeda & Silva, 2019 consists of only two genera, Choriphyllum Serville, 1838 and Phyllotettix Hancock, 1902b, combining leaf-like Caribbean wide-nosed pygmy grasshoppers (Tetrigidae, Cladonotinae). The genus Choriphyllum has four species, C. bahamense Perez-Gelabert & Otte, 1999 from the Bahamas (Hummingbird Cay Island), and C. sagrai Serville, 1838, C. saussurei Bolívar, 1887 and C. wallaceum Skejo, Kasalo & Yong, sp. nov. from Cuba. The gender of C. bahamensis is changed to C. bahamense in order to match the grammatical gender of the genus, which is neuter. Silva’s designation of C. sagrai, the type species of Choriphyllum, as nomen dubium in 2019 was incorrect as Serville’s drawing clearly points to this species, endemic to Cuba, so nothing about its identity is doubtful. The genus Phyllotettix is endemic to Jamaica, where four species live: P. rhombeus (Felton, 1765), P. foliatus (Hancock, 1902a), P. compressus (Thunberg, 1815) and P. plagiatus (Walker, 1871), comb. nov. of Choriphyllum plagiatum. Choriphyllum westwoodi Hancock, 1902a, syn. nov. is synonymous with Phyllotettix rhombeus and not with P. compressus. A new subgenus is established, Phyllotettix (Rhombotettix) subgen. nov. for P. (R.) plagiatus comb. nov. and P. (R.) rhombeus. A new species complex, Phyllotettix (compressus) sp. complex nov. is established for two morphologically close species, P. compressus, and P. foliatus; while another new species complex, Choriphyllum (sagrai) sp. complex nov. includes C. sagrai and C. wallaceum sp. nov. An annotated key to genera, subgenera, species groups and species is provided.

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Research Article Fri, 24 Mar 2023 10:44:03 +0200
Revisionary notes on Feuerborniella Vaillant, 1971, with the first record of the genus from the Afrotropical region (Diptera, Psychodidae) https://dez.pensoft.net/article/97465/ Deutsche Entomologische Zeitschrift 70(1): 121-127

DOI: 10.3897/dez.70.97465

Authors: Gunnar Mikalsen Kvifte, Santiago Jaume-Schinkel

Abstract: We establish a diagnosis for Feuerborniella Vaillant, 1971, based on a re-description of its type species, Feuerborniella obscura (Tonnoir, 1919) and comment on earlier diagnoses. Feuerborniella sinefurcata Kvifte & Jaume-Schinkel, sp. nov. is described, based on material from Tanzania, representing the second Afrotropical species of the genus following Psychoda morogorica Wagner & Andersen, 2007 which we treat as Feuerborniella morogorica comb. nov. We furthermore review earlier combinations, transferring Philosepedon ensiger Quate, 1996 and Philosepedon longistylus Quate, 1996 to Feuerborniella comb. nov., and briefly discuss generic limits with Quatiella Botosaneanu & Vaillant, 1970 and Nielseniella Vaillant, 1971.

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Research Article Thu, 16 Mar 2023 18:32:48 +0200
A remarkable new genus of Nevrorthidae (Neuroptera, Osmyloidea) from mid-Cretaceous Kachin amber of northern Myanmar https://dez.pensoft.net/article/98873/ Deutsche Entomologische Zeitschrift 70(1): 113-120

DOI: 10.3897/dez.70.98873

Authors: Hiroshi Nakamine, Shûhei Yamamoto, Yui Takahashi, Xingyue Liu

Abstract: A remarkable new genus and species of Nevrorthidae, Sisyroneurorthus aspoeckorum gen. et sp. nov., is described from mid-Cretaceous Kachin amber of northern Myanmar. This new species is the second Mesozoic representative of Nevrorthidae. The distal fusion of ScP and RA in the forewing of this new genus is shared by most Sisyridae. Given the probable sister-group relationship between Nevrorthidae and Sisyridae, our study sheds light on the evolution and morphological diversity of Nevrorthidae in the Mesozoic era.

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Research Article Thu, 16 Mar 2023 18:30:20 +0200
Revision of the genus Eotrechus Kirkaldy (Hemiptera, Heteroptera, Gerridae), with descriptions of six new species https://dez.pensoft.net/article/97117/ Deutsche Entomologische Zeitschrift 70(1): 69-111

DOI: 10.3897/dez.70.97117

Authors: Anh Duc Tran, Herbert Zettel, Robert W. Sites

Abstract: Species of the Asian genus Eotrechus Kirkaldy, 1902 (Hemiptera, Heteroptera, Gerromorpha, Gerridae, Eotrechinae) are atypical water striders because they live on the near-vertical rock surfaces of waterfalls. The taxonomy of the genus is revised and six new species are described: E. steineri sp. nov., E. kerberos sp. nov., E. thai sp. nov., E. konkakinh sp. nov., E. boukali sp. nov., and E. anderseni sp. nov. Supplementary descriptions of the apterous female and macropterous male of E. fuscus Basu, Chandra & Venkatesan, 2017 are provided. Additional and updated distribution data of species of Eotrechus are reported. Eotrechus fuscus is reported from Myanmar for the first time; E. brevipes is reported from Nepal for the first time; E. hygropetricus Andersen, 1982 is reported from southern Thailand for the first time, which extends the southernmost limit of the known distribution of the genus. A revised key to the species of Eotrechus is also provided.

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Research Article Thu, 16 Mar 2023 18:28:40 +0200
Mouthpart palp sensilla of basal Trichoptera families https://dez.pensoft.net/article/98752/ Deutsche Entomologische Zeitschrift 70(1): 55-68

DOI: 10.3897/dez.70.98752

Authors: Kseniia T. Abu Diiak, Vladimir D. Ivanov, Stanislav I. Melnitsky, Mikhail Yu. Valuyskiy, Alexandra A. Puyto

Abstract: A comparative SEM study of palp sensory surfaces in 25 caddisfly species representing seven families reveals seven types of sensilla: long trichoid, blunt chaetoid, campaniform, mushroom-like pseudoplacoid, petaloid, thick basiconic and thin basiconic sensilla. Long trichoid and chaetoid sensilla are present on all segments of both pairs of palps. First and second segments of maxillary palps bear groups of long and sclerotised chaetoid sensilla on their medial surface. Other segments of maxillary palps and all segments of labial palps have shorter and thinner chaetoid sensilla mainly on their ventromedial surfaces. Campaniform sensilla usually occur on the first segment of labial palps and second segment of maxillary palps. Mushroom-like pseudoplacoid sensilla may occupy all palp segments or only distal ones. Petaloid sensilla form sensory fields on apical segments of both pairs of palps in most studied species. Thick basiconic sensilla occur only in apical sensory complexes on tips of maxillary and labial palps. A comparison with the Lepidoptera suggests the similarity in palp sensilla and conservative evolution of the palp surface. The reconstructed ground plan for the palp sensory surfaces in Trichoptera and Amphiesmenoptera is provided.

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Research Article Thu, 16 Mar 2023 18:25:17 +0200
Revision of the South African leafhopper genus Hadroca Theron (Hemiptera, Auchenorrhyncha, Cicadellidae, Bonaspeiini) https://dez.pensoft.net/article/90851/ Deutsche Entomologische Zeitschrift 70(1): 13-54

DOI: 10.3897/dez.70.90851

Authors: Michael Stiller

Abstract: The leafhopper genus Hadroca Theron, 1974 with type species Euscelis ramosa Naudé, 1926 is re-defined here by examination of long series of specimens with varied color and wing polymorphic males and females. Five new species are added: Hadroca alacaudella sp. nov., H. alavittata sp. nov., H. bualacauda sp. nov., H. hapsistylis sp. nov. and H. treichroa sp. nov. The genus is known only from South Africa, mostly from the Fynbos Biome. All species have the aedeagus C-shaped and uniformly sclerotized, the shaft gradually tapered, commonly edentate, rarely with single, subapical, anterior process and the style with distal apophysis curved ventrad. A key to species is provided, with discussion on the differentiation from similar genera such as Bloemia Theron, 1974, Basutoia Linnavuori, 1961 and Tzitzikamaia Linnavuori, 1961 and distribution maps.

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Research Article Fri, 3 Feb 2023 17:48:41 +0200
First nymph-imago association in Polyploica confirming the distribution of Euthyplociidae (Ephemeroptera) in China https://dez.pensoft.net/article/96986/ Deutsche Entomologische Zeitschrift 70(1): 1-11

DOI: 10.3897/dez.70.96986

Authors: Xuhongyi Zheng, Xinhe Qiang, Changfa Zhou

Abstract: In this study, the imagoes and subimagoes of Polyplocia orientalis Nguyen & Bae, 2003 (Ephemeroptera, Euthyplociidae) were reared from nymphs in the lab and described for the first time. Nymphal structures are also photographed and described. The imagoes of this species can be identified by having transparent wings (except costal and subcostal areas), sterna without regular markings and penis divergent in apical half and basal half fused. The K2P distance between newly-sequenced COI gene of Chinese materials and a stored Vietnamese one is 0.10. This report not only confirms the presence of the genus and family in China, but also provides the first nymph-imago association in the genus.

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Research Article Thu, 26 Jan 2023 00:00:00 +0200
Four new species of Ischnodemus (Hemiptera, Heteroptera, Blissidae) and additional records from Argentina https://dez.pensoft.net/article/94683/ Deutsche Entomologische Zeitschrift 69(2): 283-295

DOI: 10.3897/dez.69.94683

Authors: Pablo M. Dellapé, María C. Melo

Abstract: Four new species of the genus Ischnodemus Fieber, 1837 (Hemiptera, Heteroptera, Blissidae) are described from Argentina: I. nigromaculatoides sp. nov., I. formosus sp. nov., I. infernalis sp. nov. and I. correntinus sp. nov. The known distribution is extended for three previously recorded species in the country. The species Ischnodemus brusai Dellapé & Montemayor, 2009 from Peru is transferred to Patritiodemus Slater & Ahmad, 1971. Dorsal habitus photographs and detailed images of the pygophore and right paramere of the new species and a key to the Argentinean species of Ischnodemus are given.

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Research Article Thu, 15 Dec 2022 10:35:04 +0200
Integrative taxonomy of the Lauraceae-feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan https://dez.pensoft.net/article/89864/ Deutsche Entomologische Zeitschrift 69(2): 219-281

DOI: 10.3897/dez.69.89864

Authors: Jun Souma

Abstract: Stephanitis Stål, 1873 (Hemiptera, Heteroptera, Tingidae, Tinginae, Tingini) is one of the most diverse lace bug genera in East Asia with over 50 species, many of them feeding on Ericaceae and Lauraceae. The Japanese species feeding on Lauraceae have high intraspecific and poor interspecific variation, so identification of some previous distribution records (mainly S. (Norba) aperta Horváth, 1912 and closely related taxa) is doubtful. In the present study, 5,080 specimens of Lauraceae-feeding species of Stephanitis from Japan were identified to species based on morphology, and DNA barcodes of the mitochondrial cytochrome c oxidase subunit (COI) region for 53 of those individuals were also obtained. The combined morphological and molecular evidence dataset showed that 10 species of Stephanitis feed on lauraceous trees in Japan: S. (Stephanitis) ambigua Horváth, 1912, S. (Norba) aperta, S. (N.) exigua Horváth, 1912, S. (N.) hayashii sp. nov., S. (N.) hiurai Takeya, 1963, S. (N.) ishikawai sp. nov., S. (N.) mendica Horváth, 1912, S. (S.) tabidula Horváth, 1912, S. (S.) takeyai Drake & Maa, 1955 and S. (S.) tomokunii sp. nov. Additionally, a new synonymy, S. (S.) fasciicarina Takeya, 1931, syn. nov. with S. (S.) tabidula, is proposed, a key for identifying the 10 species is provided and their distribution ranges and host plant relationships are presented.

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Research Article Thu, 15 Dec 2022 10:34:53 +0200
Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta) https://dez.pensoft.net/article/84080/ Deutsche Entomologische Zeitschrift 69(2): 151-218

DOI: 10.3897/dez.69.84080

Authors: Andrew Liston, Marko Mutanen, Mikk Heidemaa, Stephan M. Blank, Niina Kiljunen, Andreas Taeger, Matti Viitasaari, Veli Vikberg, Saskia Wutke, Marko Prous

Abstract: While working on an identification guide to the sawflies of Fennoscandia, we encountered numerous taxonomic problems, for some of which we present solutions. Dicrostema Benson, 1952 is a new synonym of Phymatoceropsis Rohwer, 1916, and not congeneric with Paracharactus MacGillivray, 1908. Two species occurring in Europe are transferred to Phymatoceropsis. Dolerus aericepsellus Heidemaa and Mutanen sp. nov. and Heptamelus viitasaarii Liston, Mutanen and Prous sp. nov. are described from Finland. Abia brevicornis Leach, 1817 nom. rev. is the valid name of Abia nitens auct. nec Linnaeus, and Abia nitens (Linnaeus, 1758) is the valid name for what has recently been called Abia sericea (Linnaeus, 1767). Tenthredo haemorrhoidalis Fabricius, 1781 is treated as an unplaced species of Hymenoptera, possibly Ichneumonoidea. Calameuta variabilis (Mocsáry, 1886) is the valid name of the species recently generally called C. haemorrhoidalis. Claremontia confusa (Konow, 1886) sp. rev. and Claremontia brevicornis (Brischke, 1883) are distinct species. Dolerus coracinus (Klug, 1818) is the valid name for D. anthracinus auct. Dolerus anthracinus (Klug, 1818) is a valid species similar to D. nitens Zaddach, 1859. Dolerus coruscans Konow, 1890 sp. rev. is a valid species. Dolerus junci (Stephens, 1835) is the valid name for Dolerus cothurnatus auct. Dolerus timidus (Klug, 1818) sp. rev. is distinguished from the similar D. pratensis (Linnaeus, 1758). A neotype is designated for Astatus punctatus Klug, 1803. Lectotypes are designated for 39 nominal species. 29 species group names are new junior synonyms. We present data on some species recently collected for the first time in Finland, including first records for the Palaearctic and West Palaearctic.

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Research Article Tue, 26 Jul 2022 21:08:45 +0300
A redefinition of Umbrageocoris with new species and new combinations (Heteroptera, Lygaeoidea, Geocoridae) https://dez.pensoft.net/article/85584/ Deutsche Entomologische Zeitschrift 69(2): 139-150

DOI: 10.3897/dez.69.85584

Authors: Péter Kóbor

Abstract: The geocorine (Hemiptera: Heteroptera: Lygaeoidea: Geocoridae) true bug genus Umbrageocoris Kóbor, 2019 is redefined based on new morphological information from newly acquired specimens. Two new species are described: U. boonei sp. nov. from continental Indomalaya and U. malipatili sp. nov. from Australia; two new combinations are proposed: U. elegantulus (Distant, 1904), comb. nov., and U. woodwardi (Malipatil, 1994), comb. nov. (both transferred from Geocoris Fallén, 1814). Keys, diagnoses, and distribution data to the discussed species are provided. Hypotheses on the origin of Umbrageocoris and its relationship to other geocorine genera in the region are formulated. New country records: U. elegantulus (Papua New Guinea), U. maai maai (Thailand, Laos) and U. woodwardi (Papua New Guinea).

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Research Article Thu, 21 Jul 2022 21:04:04 +0300
Review of the genus Sericocampsomeris Betrem, 1941 (Hymenoptera, Scoliidae) from China https://dez.pensoft.net/article/83335/ Deutsche Entomologische Zeitschrift 69(2): 125-138

DOI: 10.3897/dez.69.83335

Authors: Hua-Yan Chen, Zhen Liu, Zheng Wang, Sheng-Jie Yang, Shi-Xiao Luo

Abstract: The Chinese species of the genus Sericocampsomeris Betrem, 1941 are revised, based on morphology. Four species are recognised, of which one species is described as new, i.e. Sericocampsomeris punctata Liu & Chen, sp. nov., for which we also provide molecular sequence data to supplement its identity. A key to species of Sericocampsomeris from China and a distribution map are provided. Furthermore, we identify Sericocampsomeris punctata Liu & Chen, sp. nov. as a pollinator of Elaeocarpus sp. (Elaeocarpaceae) and recognise Leucopholis rorida (Fabricius, 1801) as a host of Sericocampsomeris rubromaculata (Smith, 1855).

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Research Article Thu, 21 Jul 2022 21:03:15 +0300
165 years of Deutsche Entomologische Zeitschrift – Editorial Response to a proposed name change for our journal https://dez.pensoft.net/article/89973/ Deutsche Entomologische Zeitschrift 69(2): 123-124

DOI: 10.3897/dez.69.89973

Authors: Dominique Zimmermann, Stefanie Paß

Abstract:

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Editorial Thu, 21 Jul 2022 21:00:44 +0300
Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini) https://dez.pensoft.net/article/83840/ Deutsche Entomologische Zeitschrift 69(1): 71-122

DOI: 10.3897/dez.69.83840

Authors: Borislav Guéorguiev, David W. Wrase, Thorsten Assmann, Jan Muilwijk, Patrice Machard

Abstract: Species of the subgenus Eremoderus Jeanne, 1996, genus Platyderus Stephens, 1827, occurring in continental Africa (excluding Macaronesia) and southwest Asia, are taxonomically revised. The following new species groups and species are defined and described, “weiratheri” group: Platyderus (Eremoderus) chatzakiae, sp. nov. (type locality: Greece, Kalymnos Island, near Stimenia Village); “iranicus-vanensis” group: Platyderus (Eremoderus) felixi, sp. nov. (type locality: Iran, Chahar Mahal va Bakhtiari Province, 10 km west of Naghan Town); Platyderus (Eremoderus) iranicus, sp. nov. (type locality: Iran, Chahar Mahal va Bakhtiari Province, 7 km NE Naghan Town); Platyderus (Eremoderus) vanensis, sp. nov. (type locality: Turkey, Van Province, Gevaş Town); Platyderus (Eremoderus) vrabeci, sp. nov. (type locality: Turkey, Nemrut Daği); “lassallei” group: Platyderus (Eremoderus) lassallei, sp. nov. (type locality: Iran, Mazandaran Province, between Nur City and Lavij Village); “davatchii” group: Platyderus (Eremoderus) klapperichi, sp. nov. (type locality: Iran, Mazandaran Province, Damavand, 2000 m); “afghanistanicus” group: Platyderus (Eremoderus) afghanistanicus, sp. nov. (type locality: Afghanistan, “Habatah”); “languidus” group: Platyderus (Eremoderus) arabicus, sp. nov. (type locality: Saudi Arabia, “Hedjaz”); Platyderus (Eremoderus) brunki, sp. nov. (type locality: Republic of Yemen, Thula District, between Kaukaban and Shibam); Platyderus (Eremoderus) irakensis, sp. nov. (type locality: Iraq, Ar Rutba District, 115 km E Ar-Rutbah Town); Platyderus (Eremoderus) jordanensis, sp. nov. (type locality: Jordan, Al-Betrā’ District, Little Petra). Six previously described species — P. brunneus Karsch, P. insignitus Bedel, P. languidus Reiche & Saulcy, P. ledouxi Morvan, P. taghizadehi Morvan, and P. weiratheri Mařan — are redescribed based on type and/or non-type material. P. davatchii Morvan placed as a member of the subgenus was not treated due to the lack of material available for study. The following new nomenclature acts are proposed: Platyderus brunneus Karsch, 1881, stat. rev., is removed from synonymy with Feronia languida Reiche & Saulcy, 1855; Platyderus elegans Bedel, 1900, syn. nov., is proposed as junior synonym of Platyderus brunneus Karsch, 1881; Platyderus ferrantei Reitter, 1909 is proposed as subspecies Platyderus brunneus ferrantei Reitter, 1909, stat. nov. In order to preserve the stability of nomenclature, lectotypes are designated for: Feronia languida Reiche & Saulcy, Platyderus brunneus Karsch, and Platyderus weiratheri Mařan. Keys to identification of the male and female specimens of the species from the regions studied are provided.

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Research Article Mon, 13 Jun 2022 17:04:17 +0300
Identity of Zorotypus juninensis Engel, 2000, syn. nov. revealed: it is conspecific with Centrozoros hamiltoni (New, 1978) (Zoraptera, Spiralizoridae) https://dez.pensoft.net/article/83154/ Deutsche Entomologische Zeitschrift 69(1): 65-70

DOI: 10.3897/dez.69.83154

Authors: Petr Kočárek, Ivona Horká

Abstract: Zorotypus juninensis Engel, 2000, was previously diagnosed based on the external morphology of female and male specimens without description of the male copulatory organ, which is an important character for classification in Zoraptera. Based on a detailed morphological study of the Zorotypus juninensis Engel, 2000 type collections deposited in the American Museum of Natural History in New York, and based on the comparison with the holotype male of Centrozoros hamiltoni (New, 1978), we have determined that these two species are conspecific. We therefore formally synonymize Zorotypus juninensis Engel, 2000, syn. nov. with Centrozoros hamiltoni (New, 1978). Morphological characters and phylogenetic relationships of Centrozoros Kukalova-Peck & Peck, 1993 are also discussed in this report.

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Research Article Wed, 8 Jun 2022 10:41:03 +0300
Revision of the genus Woldstedtius Carlson, 1979 (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan https://dez.pensoft.net/article/80492/ Deutsche Entomologische Zeitschrift 69(1): 45-64

DOI: 10.3897/dez.69.80492

Authors: Shunsuke Morishita, Kyohei Watanabe

Abstract: Japanese species of the genus Woldstedtius Carlson, 1979 are revised. Nine species are recorded from Japan, including two new species, W. alpicola sp. nov. and W. punctatus sp. nov. Woldstedtius biguttatus (Gravenhorst, 1829) is newly recorded from Japan. Taxonomic status of W. flavolineatus kuroashii (Uchida, 1957) is changed from the subspecies of W. flavolineatus (Gravenhorst, 1829) to a separated species. Woldstedtius holarcticus (Diller, 1969) is newly synonymized under W. kuroashii (Uchida, 1957). A key to Japanese species of this genus is provided.

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Review Article Mon, 4 Apr 2022 14:20:05 +0300
Character analysis and descriptions of Eocene sphodrine fossils (Coleoptera, Carabidae) using light microscopy, micro-CT scanning, and 3D imaging https://dez.pensoft.net/article/79931/ Deutsche Entomologische Zeitschrift 69(1): 19-44

DOI: 10.3897/dez.69.79931

Authors: Joachim Schmidt, Stephan Scholz, Kipling Will

Abstract: Of the 12 specimens of Calathus-like sphodrine beetles presently known from Baltic and Rovno amber deposits, 11 specimens were investigated using light microscopy, micro-CT scanning, and 3D imaging techniques. For the first time, many significant diagnostic characters of the external morphology and male and female genitalia of Eocene Sphodrini were studied in detail. Based on these data, three fossil species are diagnosed and placed in a natural group characterized by a derived pattern in elytral chaetotaxy and microsculpture and therefore the genus Quasicalathus Schmidt & Will, gen. nov. is described to comprise these species. Due to the presence of a styloid right paramere, Quasicalathus gen. nov. is considered a member of the sphodrine “P clade” of Ruiz et al. (2009). However, given the absence of synapomorphies of any species group of the P clade, the systematic position of Quasicalathus gen. nov. within this clade remains unresolved. The Baltic amber species Calathus elpis Ortuño & Arillo, 2009 is redescribed based on additional, fossil, non-holotype material and transferred to Quasicalathus gen. nov. Identification of the additional C. elpis fossil material remains slightly uncertain due to the non-availability of the holotype for direct comparison coupled with doubts regarding the accuracy of certain character states presented in its original description. Two species are newly described: Quasicalathus agonicollis Schmidt & Will, sp. nov., from Baltic amber, and Q. conservans Schmidt & Will, sp. nov., from Rovno amber.

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Research Article Wed, 9 Feb 2022 11:11:21 +0200
Phylogeny and age of cockroaches: a reanalysis of mitogenomes with selective fossil calibrations https://dez.pensoft.net/article/68373/ Deutsche Entomologische Zeitschrift 69(1): 1-18

DOI: 10.3897/dez.69.68373

Authors: Xin-Ran Li

Abstract: In spite of big data and new techniques, the phylogeny and timing of cockroaches remain in dispute. Apart from sequencing more species, an alternative way to improve the phylogenetic inference and time estimation is to improve the quality of data, calibrations and analytical procedure. This study emphasizes the completeness of data, the reliability of genes (judged via alignment ambiguity and substitution saturation), and the justification for fossil calibrations. Based on published mitochondrial genomes, the Bayesian phylogeny of cockroaches and termites is recovered as: Corydiinae + (((Cryptocercidae + Isoptera) + ((Anaplectidae + Lamproblattidae) + (Tryonicidae + Blattidae))) + (Pseudophyllodromiinae + (Ectobiinae + (Blattellinae + Blaberidae)))). With two fossil calibrations, namely, Valditermes brenanae and Piniblattella yixianensis, this study dates the crown Dictyoptera to early Jurassic, and crown Blattodea to middle Jurassic. Using the ambiguous ‘roachoid’ fossils to calibrate Dictyoptera+sister pushes these times back to Permian and Triassic. This study also shows that appropriate fossil calibrations are rarer than considered in previous studies.

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Research Article Fri, 7 Jan 2022 13:24:28 +0200
New species of Omma Newman from mid-Cretaceous Burmese amber (Coleoptera, Archostemata, Ommatidae) https://dez.pensoft.net/article/74174/ Deutsche Entomologische Zeitschrift 68(2): 341-348

DOI: 10.3897/dez.68.74174

Authors: Yan-Da Li, Di-Ying Huang, Chen‑Yang Cai

Abstract: A new fossil species of the extant archostematan genus Omma Newman, O. forte sp. nov., is reported from mid-Cretaceous amber from northern Myanmar. The extinct ommatid genus, Cionocups Kirejtshuk from the same deposit, is considered as a junior synonym of Omma, and C. manukyani is transferred to Omma, as O. manukyani comb. nov. A key to species of Omma from Burmese amber is also provided.

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Research Article Tue, 30 Nov 2021 12:23:58 +0200
Revision of Indian species of Phanuromyia Dodd, 1914 (Platygastroidea, Scelionidae) with descriptions of new species https://dez.pensoft.net/article/70497/ Deutsche Entomologische Zeitschrift 68(2): 309-339

DOI: 10.3897/dez.68.70497

Authors: Kamalanathan Veenakumari, Sreedevi Kolla, Prashanth Mohanraj, Farmanur Rahman Khan

Abstract: The species of the genus Phanuromyia Dodd, 1914, belonging to the subfamily Telenominae are egg parasitoids of planthoppers belonging to the families Issidae, Flatidae, Fulgoridae and Ricaniidae (Hemiptera: Fulguroidea). So far eleven species of Phanuromyia are known from India. Fifteen new species of Phanuromyia Dodd are described here: P. chalukya sp. nov., P. chera sp. nov., P. chola sp. nov., P. ganga sp. nov., P. hoysala sp. nov., P. kadamba sp. nov., P. kakatiya sp. nov., P. kanva sp. nov., P. nirvighna sp. nov., P. pallava sp. nov., P. pandya sp. nov., P. rashtrakuta sp. nov., P. satavahana sp. nov., P. tuluva sp. nov. and P. vakataka sp. nov. A key to females of all the Indian species of Phanuromyia is provided.

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Research Article Tue, 30 Nov 2021 12:23:10 +0200
Limnomma, a new genus of Ommatidae from the Middle Jurassic Daohugou beds (Coleoptera, Archostemata) https://dez.pensoft.net/article/71880/ Deutsche Entomologische Zeitschrift 68(2): 299-308

DOI: 10.3897/dez.68.71880

Authors: Yan-Da Li, Erik Tihelka, Hong Pang, Di-Ying Huang, Chen‑Yang Cai

Abstract: The relictual archostematan beetle family Ommatidae attained high diversity during the Mesozoic. Despite their once high taxonomic diversity and morphological disparity, many Mesozoic ommatid taxa remain poorly understood, partly due to limited preservation. Here we report an exceptionally well-preserved fossil, which we describe as a new ommatid genus and species, Limnomma daohugouense gen. et sp. nov., from the mid-Jurassic Daohugou Lagerstätte in Northeast China. The new genus can be most easily distinguished from other ommatids by the presence of a circular non-tuberculate region on ventrite 5. The new taxon is discussed in relation to the classification of the Mesozoic genera Brochocoleus and Burmocoleus.

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Research Article Fri, 3 Sep 2021 17:46:45 +0300
Revision of the genus Xanthopimpla Saussure, 1892 (Hymenoptera, Ichneumonidae, Pimplinae) from Japan https://dez.pensoft.net/article/69768/ Deutsche Entomologische Zeitschrift 68(2): 269-297

DOI: 10.3897/dez.68.69768

Authors: Kyohei Watanabe, Rikio Matsumoto

Abstract: Japanese species of the genus Xanthopimpla Saussure, 1892 are revised. A total of 15 species are recorded from Japan, including three new species, X. nipponensis sp. nov. X. sylvicola sp. nov. and X. yoshimurai sp. nov. and three species, X. honorata (Cameron, 1899), X. minuta Cameron, 1905 and X. trias Townes & Chiu, 1970, newly recorded from Japan. Host, habitat, overwintering and distribution patterns of Japanese Xanthopimpla species are discussed.

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Research Article Wed, 1 Sep 2021 17:46:05 +0300
Where did the Central European populations of Ornatoraphidia flavilabris (Costa) come from? (Neuropterida, Raphidioptera, Raphidiidae) https://dez.pensoft.net/article/70814/ Deutsche Entomologische Zeitschrift 68(2): 249-259

DOI: 10.3897/dez.68.70814

Authors: Horst Aspöck, Ulrike Aspöck, Julia Walochnik, Edwin Kniha

Abstract: Ornatoraphidia flavilabris (Costa, 1851) is one of 15 snakefly species occurring in southern parts of Central Europe. It is a polycentric Mediterranean faunal element with refugia in the Apennine Peninsula and the Balkan Peninsula. Two phylogeographic questions are dealt with in this paper: (1) Is it possible to differentiate, morphologically or genetically, the Balkanic populations from the Italian? (2) Did the species reach Central Europe from the Balkan or Apennine Peninsula? These questions were investigated using morphological and molecular biological methods. No morphological characters were uncovered which could serve to differentiate specimens from either distribution center. However, differences were detected in cox1, cox3 and 28S genes which allow for a reliable differentiation. Central European populations were largely identical with populations from Italy, but distinctly different from specimens from Greece. This could lead one to assume that the species migrated from Italy to Central Europe, although colonization from the southeast would appear easier due to more favorable orographic conditions. This discrepancy may be explained by the apparent absence of O. flavilabris from the large central part of the Balkan Peninsula, so that a gap exists between the southern and northern areas inhabited by O. flavilabris. Moreover, the species does not occur in eastern parts of Europe. Thus it would be more probable to assume that the occurrence of the species in the northwest Balkan Peninsula can be traced to migrations from the Apennine Peninsula to areas north and northeast of the Adriatic Sea, where O. flavilabris may have colonized the southeast of Central Europe. A migration of Adriatomediterranean faunal elements from the northwest Balkan Peninsula to Central Europe might be of more significance than previously assumed.

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Research Article Mon, 30 Aug 2021 16:47:08 +0300
Revision of Iranian Schoenlandella Cameron, 1905 (Hymenoptera, Braconidae, Cardiochilinae) with descriptions of two new species from Hormozgan province https://dez.pensoft.net/article/69090/ Deutsche Entomologische Zeitschrift 68(2): 261-268

DOI: 10.3897/dez.68.69090

Authors: Ilgoo Kang, Ali Ameri, Michael J. Sharkey

Abstract: The Iranian species of the genus Schoenlandella Cameron, 1905 are revised based on morphological characters. The current work provides a key to species of Iranian Schoenlandella and species descriptions of two new species: S. angustigena Kang sp. nov. and S. latigena Kang sp. nov. This work results in increasing the number of Iranian Schoenlandella species from one to three species.

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Research Article Mon, 30 Aug 2021 12:43:58 +0300
The Pyrenean species of Chelidura (Dermaptera, Forficulidae) https://dez.pensoft.net/article/68020/ Deutsche Entomologische Zeitschrift 68(2): 235-248

DOI: 10.3897/dez.68.68020

Authors: Pilar Jurado-Angulo, Yolanda Jiménez-Ruiz, Mario García-París

Abstract: The Pyrenees are inhabited by scattered populations of earwigs of the genus Chelidura Latreille, 1825. There is some controversy about the specific assignment of these populations: while most authors assign them to C. pyrenaica (Gené, 1832), other consider that C. aptera (Mégerlé, 1825) is also present in the Pyrenees. The main objective of this work was to revise the identity and synonyms of Pyrenean Chelidura. Specimens from recent fieldwork and collections (MNCN-CSIC) were used for morphological and molecular studies (cytochrome oxidase 1). All Pyrenean specimens shared similar cox1 sequences, very divergent from those of Alpine C. aptera. As a consequence, the variability observed in male cerci morphology from the Pyrenees, ranging from long and slightly curved to short and very curved, corresponded to C. pyrenaica, and the presence of C. aptera in the Pyrenees can be rejected. As previously suggested by Maccagno (1933) and Fontana et al. (2021), the revision of the synonymic list uncovered the misplacement of the name F. simplex Germar, 1825 under the synonymy of C. aptera, while it rather represents a synonym of C. pyrenaica (syn. nov.). Forficula simplex has nomenclatural priority over C. pyrenaica, however both names meet the requirements of the article 23.9.1 of the International Code of Zoological Nomenclature to retain the prevailing usage of C. pyrenaica (nomen protectum) over F. simplex (nomen oblitum). Additionally, we discuss the taxonomic status of Chelidura arverna David & Van Herrewege, 1973 stat. nov. from the French Massif Central.

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Research Article Tue, 10 Aug 2021 09:53:22 +0300
A third species of the rare ant genus Rotastruma (Hymenoptera, Formicidae) from Cambodia, with illustrated generic diagnoses and key to species https://dez.pensoft.net/article/63478/ Deutsche Entomologische Zeitschrift 68(2): 225-233

DOI: 10.3897/dez.68.63478

Authors: Shingo Hosoishi, Seiki Yamane, Heng Sokh

Abstract: The myrmicine genus Rotastruma is a rarely collected arboreal ant group and only two species, R. recava Bolton and R. stenoceps Bolton, are known from the Oriental Region. A new species, R. epispina sp. nov., is described from Cambodia based on the worker and queen castes. The new species is distinguished from R. recava and R. stenoceps by having the propodeal spine directed upwards and elongated peduncle of the petiole. Herein, we provide an update to the identification key and diagnostic characters for the genus Rotastruma.

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Research Article Mon, 5 Jul 2021 08:45:54 +0300
Balticeler kerneggeri gen. nov., sp. nov., an enigmatic Baltic amber fossil of the ground beetle subfamily Trechinae (Coleoptera, Carabidae) https://dez.pensoft.net/article/66181/ Deutsche Entomologische Zeitschrift 68(1): 207-224

DOI: 10.3897/dez.68.66181

Authors: Joachim Schmidt, Stephan Scholz, David R. Maddison

Abstract: Balticeler kerneggeri gen. nov., sp. nov., is described based on six fossil specimens preserved in Eocene Baltic amber and imaged using light microscopy and X-ray micro-computed tomography. Based on certain characters observed in the fossil species it is considered a “middle grade” Carabidae, outside of the large family Harpalinae (as it possesses a scrobal seta, the lack of which is a synapomorphy of that subfamily), but possessing four synapomorphies that indicate Balticeler belongs to a large clade of carabids including Harpalinae (anisochaetous Grade B antennal cleaner, conjunct mesocoxae, closed procoxal cavities, and a well-developed external lobe of the metepimeron). This remarkable beetle has several striking features, including lack of externally-visible sexually dimorphic characters, lack of lateral borders on the pronotum, and very long and thin mandibles and maxillae. In combination, these states are unique within Carabidae. We consider the presence of a dorsally completely open aedeagal median lobe as a synapomorphy of the fossil species with the subfamily Trechinae, a pubescent and relatively long second antennomere and a 4+2+2 pattern of umbilicate setae as synapomorphies of the supertribe Trechitae, and a quadrisetose clypeus as a synapomorphy with the Trechitae clade Bembidarenini + Trechini sensu Maddison et al. (2019). As it lacks a synapomorphy of Bembidarenini + Trechini, we propose that it is a member of the stem group of that clade.

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Research Article Wed, 2 Jun 2021 08:55:53 +0300
Four new species of Agraphydrus Régimbart, 1903 with additional faunistic record from China (Coleoptera, Hydrophilidae, Acidocerinae) https://dez.pensoft.net/article/66200/ Deutsche Entomologische Zeitschrift 68(1): 189-205

DOI: 10.3897/dez.68.66200

Authors: Zhen-ming Yang, Fenglong Jia, Lu Jiang, Qiang Guo

Abstract: Four new species of Agraphydrus Régimbart are described from China: A. pseudoniger sp. nov. from Shangyou County, Jiangxi Province, A. komareki sp. nov. from Shangchuan island, Taishan County, Guangdong Province, A. sabulosus sp. nov. from Fengkai, Guangdong Province, A. dapengensis sp. nov. from Dapeng peninsula, Shenzhen, Guangdong Province. Diagnosis and illustration of the new species are provided. The key given by Komarek and Hebauer (2018) to Chinese species of Agraphydrus Régimbart is updated.

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Research Article Tue, 18 May 2021 15:32:18 +0300
First instar nymphs of two peltoperlid stoneflies (Insecta, Plecoptera, Peltoperlidae) https://dez.pensoft.net/article/65540/ Deutsche Entomologische Zeitschrift 68(1): 179-188

DOI: 10.3897/dez.68.65540

Authors: Shodo Mtow, Tadaaki Tsutsumi

Abstract: The first instar nymphs of two peltoperlid stoneflies, i.e., Microperla brevicauda Kawai, 1958 of Microperlinae and Yoraperla uenoi (Kohno, 1946) of Peltoperlinae, were examined and described. Additionally, the phylogeny and groundplan of the first instar nymphs of Peltoperlidae and Plecoptera were considered. The first instar nymphs of M. brevicauda have a slender body with a prognathous head of typical shape; they represent a groundplan in Plecoptera. On the other hand, the first instar nymphs of Y. uenoi have a broad, cockroach-like body with an orthognathous and shortened head, the latter being regarded as a potential autapomorphy of Peltoperlinae. Such differences in body shape between the subfamilies are speculated to arise from heterochrony. The three-segmented cerci of Y. uenoi are characteristic to Systellognatha, whereas the four-segmented cerci of M. brevicauda were independently acquired within Microperlinae. The structure and distribution pattern of chloride cells in the first instar nymphs of Plecoptera were also discussed. The presence of coniform chloride cells is a potential groundplan of Arctoperlaria. One to two pairs of chloride cells are distributed on the first nine abdominal segments of M. brevicauda; this represents a groundplan character of Systellognatha. On the other hand, one to four pairs of chloride cells are found on the second to ninth abdominal segments of Y. uenoi; this distribution pattern may be an apomorphic groundplan of Peltoperlinae.

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Research Article Fri, 7 May 2021 16:55:56 +0300
Kazukuru gen. nov. – a new Ricaniidae planthopper from Solomon Islands (Hemiptera, Fulgoromorpha) https://dez.pensoft.net/article/63635/ Deutsche Entomologische Zeitschrift 68(1): 165-177

DOI: 10.3897/dez.68.63635

Authors: Adam Stroiński

Abstract: A new monotypic genus of ricaniid planthoppers (Hemiptera: Fulgoromorpha: Ricaniidae) from New Georgia Island (Solomon Islands), Kazukuru gen. nov., is described for K. zingiberis sp. nov. (type species). Habitus, female, external and internal genital structures of the new species are described and illustrated.

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Research Article Wed, 31 Mar 2021 15:58:03 +0300
Records and descriptions of caddisflies from Natma Taung National Park and adjacent localities in the Chin Hills of Myanmar (Insecta, Trichoptera) https://dez.pensoft.net/article/61819/ Deutsche Entomologische Zeitschrift 68(1): 139-164

DOI: 10.3897/dez.68.61819

Authors: Wolfram Mey, Hans Malicky

Abstract: During several excursions to the Chin Hills of Myanmar from 2001 to 2015, rich collections of caddisflies were made which form the basis of the present taxonomic and biogeographic study. A total of 106 species were identified including seven new species. They are described as Arctopsyche subflava sp. nov., Hydromanicus abdominalis sp. nov., Cheumatopsyche janosolahi sp. nov., Lepidostoma subpanaitos sp. nov., Aplatyphylax pumilus sp. nov., Adicella natmataungensis sp. nov. and Triaenodes mindatensis sp. nov. Illustrations of the male genitalia and images of the adults are provided. Two species names were recognised as junior synonyms: Hydropsyche athamas Malicky & Chantaramongkol, 2000, = Hydropsyche januha Oláh & Barnard, 2008, syn. nov.; Hydropsyche khasigiri Oláh & Barnard, 2008, = Hydropsyche kiogupa Oláh & Schefter, 2008, syn. nov. In an attempt to determine the biogeographic character of the fauna, the known ranges of all resident species were plotted into three longitudinal transects from 85°–95°E, 95–98°E and 98°–108°E, ranging from the eastern Himalayas to northern Vietnam. About half of the species have ranges extending over all three transects. The fauna is equally composed of western and eastern species qualifying the Chin Hills as part of a transition zone including palearctic elements. Most of the species, which are widely distributed in south-east Asia, are members of the suborder Annulipalpia. The investigations of the authors yielded 77 autochthonous species, which have been unrecorded from the country including the newly-described taxa. This brings the number of species records from Mynamar to 304.

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Research Article Fri, 26 Mar 2021 10:19:34 +0200
Morphology, pollen preferences and DNA-barcoding of five Austrian species in the Colletes succinctus group (Hymenoptera, Apidae) https://dez.pensoft.net/article/55732/ Deutsche Entomologische Zeitschrift 68(1): 101-138

DOI: 10.3897/dez.68.55732

Authors: Katharina Zenz, Herbert Zettel, Michael Kuhlmann, Harald W. Krenn

Abstract: Most species of the Colletes succinctus group sensu Noskiewicz, 1936 are taxonomically uncertain. This study has chosen an integrative approach, including pollen analysis, morphology, male genitalia, morphometry, cuticle sculpture and DNA-barcoding (CO1) to investigate the five species that were reported from Austria. It includes a detailed analysis of the male genitalia and the first description of the C. pannonicus male. A syntype male from the island of Crete was designated as the lectotype of Colletes succinctus brevigena Noskiewicz, 1936 to fix the species identity. New distinguishing characters were found: in females the shape of the dorsal end of the fovea facialis and, in both sexes, the structure of maxillary palpi, as well as the different puncturation on the mesopleura. Unknown structures on sterna and genitalia of the males proved to be reliable morphological characters. An identification key is provided for all studied species. Morphometry of females did not allow a clear distinction of species. CO1 sequencing confirmed previous studies that only C. collaris clearly deviates from the other species, including C. pannonicus that was analysed for the first time. Pollen analysis showed polylectic, as well as oligolectic, pollen-collecting behaviour. The collected pollen of C. pannonicus confirmed the field observations that this species is strictly oligolectic on Tripolium pannonicum. Due to pronounced intraspecific variation, it is assumed that the species of the C. succinctus group are either species in statu nascendi or very young species. Therefore, it remains important to include ecological data in species identification.

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Research Article Tue, 2 Feb 2021 16:14:43 +0200
A new fossil species of the genus Bibio, with an update on bibionid flies from Baltic and Rovno amber (Diptera, Bibionidae) https://dez.pensoft.net/article/60611/ Deutsche Entomologische Zeitschrift 68(1): 81-99

DOI: 10.3897/dez.68.60611

Authors: John Skartveit

Abstract: Species of Bibionidae from Baltic amber are reevaluated based on newly discovered material, and a key to the species is given. Bibio succineus sp. nov. is described based on one male specimen, this is the first Bibio named from Baltic amber. The males of Hesperinus electrus Skartveit, 2009 and Penthetria montanaregis Skartveit, 2009 are redescribed. A single, autoclave treated specimen of Penthetria sp. is described but not formally named. Plecia tenuicornis Skartveit, 2009 is found to be a synonym of Plecia hoffeinsorum Skartveit, 2009, this species is recorded for the first time from Rovno amber, and both sexes of the species redescribed. Additional specimens of Plecia clavifemur Skartveit, 2009 and Dilophus crassicornis Skartveit, 2009 are described. Two female specimens probably belonging to the species discussed as Dilophus sp. by Skartveit (2009) are described, but not formally named.

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Research Article Wed, 27 Jan 2021 07:37:11 +0200
Biogeographical and evolutionary aspects of a Guineo-Congolian bushcricket tribe: Revision of the genera Cestromoecha Karsch, 1893 and Poreuomena Brunner von Wattenwyl, 1878, with the description of new species (Orthoptera, Tettigoniidae, Phaneropterinae) https://dez.pensoft.net/article/60193/ Deutsche Entomologische Zeitschrift 68(1): 45-79

DOI: 10.3897/dez.68.60193

Authors: Claudia Hemp, Bruno Massa

Abstract: The genera Cestromoecha and Poreuomena of the tribe Poreuomenini in Phaneropterinae are revised and new generic characters are given for both genera, and six new species are described in Poreuomena. The newly described species are P. biaculeata sp. nov., P. eala sp. nov., P. gracilicercata sp. nov., P. ivoriana sp. nov., P. matthaei sp. nov., and P. tshuapa sp. nov. Based on characters defining the two genera, three species so far listed under Cestromoecha are transferred to Poreuomena: P. crassipes Karsch, 1890, P. laeglae (Massa, 2015), and P. magnicerca (Massa, 2013). One species of Cestromoecha, C. mundamensis Karsch, 1896, is synonymised with C. tenuipes (Karsch, 1890) since no morphological differences were detected between the type specimens. Thus, two species remain with Cestromoecha, and Poreuomena now contains 16 species. Morphological closely-related species of Poreuomena suggest rapid speciation in the Congo Basin due to several expansions and shrinkages of the Guineo-Congolian forest belt since the Oligocene. At least two different morphological lineages are discernible. On the other hand the genus Cestromoecha Karsch, 1893 is a species-poor taxon.

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Research Article Tue, 5 Jan 2021 16:49:00 +0200
Review of the world species of Paroplitis Mason, 1981 (Hymenoptera, Braconidae, Microgastrinae), with description of three new species https://dez.pensoft.net/article/59641/ Deutsche Entomologische Zeitschrift 68(1): 33-43

DOI: 10.3897/dez.68.59641

Authors: Shunpei Fujie, George Japoshvili, Jose Fernandez-Triana

Abstract: The world species of the microgastrine genus Paroplitis (Hymenoptera: Braconidae) are revised. Three new species are described, P. horticola Fujie & Fernandez-Triana, sp. nov. and P. japonicus Fujie & Fernandez-Triana, sp. nov. from Japan and P. kakhetiensis Fujie, Japoshvili & Fernandez-Triana, sp. nov. from Georgia. P. vietnamensis van Achterberg & Fernandez-Triana, 2013 is re-described, based on additional specimens. P. wesmaeli Ruthe, 1860 is recorded from Georgia for the first time. A key to the nine known species (eight described and one undescribed) of the genus is provided.

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Research Article Tue, 5 Jan 2021 14:34:54 +0200
Comparative morphology of the musculature of the sting apparatus in Ampulex compressa (Hymenoptera, Ampulicidae) and Sceliphron destillatorium (Hymenoptera, Sphecidae) https://dez.pensoft.net/article/58217/ Deutsche Entomologische Zeitschrift 68(1): 21-32

DOI: 10.3897/dez.68.58217

Authors: Stefan Graf, Maraike Willsch, Michael Ohl

Abstract: The sting apparatus of aculeate Hymenoptera is derived from the ovipositor and is their most prominent apomorphy. In contrast to the frequently analysed sclerites of the sting apparatus, the associated musculature has largely been neglected. In this study, we use micro-computed tomography to present a detailed description of the musculature of the sting apparatus of Ampulex compressa (Ampulicidae) and Sceliphron destillatorium (Sphecidae). We found that 12 of 15 muscles corresponding to the sting apparatus are homologous between both species examined and 13 muscles in comparison with Hymenoptera described in the literature. All muscles identified as critical for the act of stinging were found in both species. Moreover, we found the ventral tergum 8-tergum 9 muscle and the tergum 8-tergum 8 muscles in A. compressa and the second valvifer-second valvifer muscle in S. destillatorium. For the first time, we describe the ventral tergum 8-tergum 9 muscle and the second valvifer-second valvifer muscle that interconnects both body sides, in Hymenoptera.

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Research Article Tue, 5 Jan 2021 13:41:00 +0200
Philibaetis gen. nov., a new genus from the Philippines (Ephemeroptera, Baetidae) https://dez.pensoft.net/article/59462/ Deutsche Entomologische Zeitschrift 68(1): 1-20

DOI: 10.3897/dez.68.59462

Authors: Thomas Kaltenbach, Jhoana M. Garces, Jean-Luc Gattolliat

Abstract: Investigations of type material and new material from the Philippines (Luzon) revealed that Baetis luzonensis Müller-Liebenau, 1982 and B. realonae Müller-Liebenau, 1982 do not belong to Baetis Leach, 1815. A new genus, Philibaetis gen. nov., is described to accommodate both species and both are re-described based on larvae. The new genus is characterised by having a rectangular labrum with a submarginal row of long, simple setae on the dorsal surface and ventrally on lateral margins long, simple, spine-like setae, on anterolateral margins long, feathered setae and medially long, bifid setae and a partial, submarginal row of lanceolate setae. Both mandibles have blade-like incisors and dorsally, a mediolateral patch of long, spine-like setae; additionally, the left mandible has a tuft of long, partly branched setae at the base of the subtriangular process. Philibaetis gen. nov. is further characterised by a hypopharynx with a medial tuft of stout setae and anterolaterally, two smaller tufts of stout setae, a galea-lacinia with the distal denti-seta tooth-like and directed against canines, a fore femur apically with stout setae, both on anterior and posterior side and without a femoral patch and a claw with one row of denticles and two or three subapical setae. The protogonostyli under the cuticle of male last instar larvae are folded in the Labiobaetis type, excluding their affiliation to the genus Baetis. COI sequences were obtained from both species. The genetic distance (Kimura 2-parameter) between them is 17.5% on average. Very limited genetic distances of 0% to 3% (0.75% on average) were found between specimens of P. luzonensis comb. nov.

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Research Article Tue, 5 Jan 2021 12:19:29 +0200
The subgenera Glabrobracon Fahringer, Lucobracon Fahringer and Uncobracon Papp of the genus Bracon Fabricius (Hymenoptera, Braconidae, Braconinae) in China, with the description of eleven new species https://dez.pensoft.net/article/57668/ Deutsche Entomologische Zeitschrift 67(2): 209-252

DOI: 10.3897/dez.67.57668

Authors: Yang Li, Jun-hua He, Xue-xin Chen

Abstract: The species of three subgenera of the genus Bracon Fabricius (Hymenoptera, Braconidae, Braconinae), i.e. Glabrobracon Fahringer, 1927; Lucobracon Fahringer, 1927; and Uncobracon Papp, 1996 from China are revised and 31 species are recognised, including 11 new species, i.e. Bracon (Glabrobracon) indistinctus sp. nov., B. (G.) leptotes sp. nov., B. (G.) longistriatus sp. nov., B. (G.) megaventris sp. nov., B. (Lucobracon) brevicarinatus sp. nov., B. (L.) coarctatus sp. nov., B. (L.) curculiovorus sp. nov., B. (L.) flavitestaceus sp. nov., B. (L.) quadratus sp. nov., B. (Uncobracon) eurysulcatus sp. nov. and B. (U.) longwangshanensis sp. nov. The new species are described and illustrated. Keys to the subgenera of the genus Bracon and the Chinese species of three subgenera Glabrobracon, Lucobracon and Uncobracon are provided.

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Research Article Thu, 19 Nov 2020 08:38:05 +0200
A DNA barcode library for ground beetles of Germany: the genus Agonum Bonelli, 1810 (Insecta, Coleoptera, Carabidae) https://dez.pensoft.net/article/56163/ Deutsche Entomologische Zeitschrift 67(2): 197-207

DOI: 10.3897/dez.67.56163

Authors: Michael J. Raupach, Karsten Hannig, Jérome Morinière, Lars Hendrich

Abstract: The ground beetle genus Agonum Bonelli, 1810 is a large genus of the tribe Platynini with many species that show high amounts of intraspecific variations, making a correct identification challenging. As part of the German Barcode of Life initiative, this publication provides a comprehensive DNA barcode library for species of Agonum that are reported for Germany. In total, DNA barcodes from 258 beetles and 23 species were analysed using the Barcode of Life Data System (BOLD) workbench, including sequences from former studies and 68 newly-generated sequences. The neighbour-joining analyses, based on K2P distances, revealed distinct clustering for all studied species, with unique Barcode Index Numbers (BINs) for 15 species (65%). BIN sharing but distinct clustering was found for three species pairs: Agonum micans/Agonum scitulum, Agonum impressum/Agonum sexpunctatum and Agonum duftschmidi/Agonum emarginatum. The given dataset and its analysis represent another important step in generating a comprehensive DNA barcode library for the ground beetles of Germany and Central Europe in terms of modern biodiversity research.

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Research Article Mon, 19 Oct 2020 17:00:53 +0300
Revision of the Plagiolepis schmitzii group with description of Pl. invadens sp. nov. – a new invasive supercolonial species (Hymenoptera: Formicidae) https://dez.pensoft.net/article/53199/ Deutsche Entomologische Zeitschrift 67(2): 183-196

DOI: 10.3897/dez.67.53199

Authors: Bernhard Seifert

Abstract: Using high-resolution stereomicroscopy and exploratory data analyses, a taxonomic revision of the cryptic species close to Plagiolepis schmitzii Forel, 1895, called Pl. schmitzii group, was conducted. Morphology was numerically recorded under highly standardised conditions considering absolute size and 16 shape, pubescence and surface characters. A key to the non-parasitic Westpalaearctic species of the ant genus Plagiolepis Mayr, 1861 is provided which firstly separates, on species group level, the Pl. pygmaea (Latreille) species group, the Pl. pallescens Forel species group and the Pl. schmitzii species group and, finally, on species level, the cryptic species of the latter group. The recognised species of the Pl. schmitzii species group are Pl. schmitzii Forel, 1895 (invasive species), Pl. barbara Santschi, 1911, Pl. atlantis Santschi, 1920 and Pl. invadens sp. nov. (invasive species) that is described as new from a supercolony in Germany. Based on morphological arguments, the taxa Pl. barbara var. madeirensis Emery, 1921, Pl. maura polygyna Santschi, 1922 and Pl. schmitzii var. tingitana Santschi, 1936 are recognised as junior synonyms of Pl. schmitzii, the taxa Pl. schmitzii crosi Santschi, 1920, Pl. pallescens var. kabyla Santschi, 1920 and Pl. perperamus Salata et al., 2018 as junior synonyms of Pl. atlantis and the taxon Pl. maura Santschi, 1920 as junior synonym of Pl. barbara. A concluding comparative section suggests that pre-adaptations for anthropogenous dispersal and transformation to supercoloniality in introduction areas are apparently common traits in Plagiolepis ants.

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Research Article Mon, 21 Sep 2020 15:16:35 +0300
Phylogeny and classification of the genus-group taxa of Loxandrina (Coleoptera, Carabidae, Abacetini) https://dez.pensoft.net/article/55985/ Deutsche Entomologische Zeitschrift 67(2): 151-182

DOI: 10.3897/dez.67.55985

Authors: Kipling Will

Abstract: Bayesian and parsimony phylogenetic analyses of combined and partitioned datasets of molecular (partial sequences of 28S, wg, COI, and CAD) and morphological (51 characters of adults) data for exemplar taxa of five outgroup and 76 ingroup abacetine carabids resulted in a monophyletic Loxandrina Erwin & Sims, 1984 that is split into Australian and American clades. The genus Loxandrus LeConte, 1853 as previously delimited is not monophyletic relative to numerous genus-level taxa in Abacetini Chaudoir, 1873 and is restricted to a subgenus of North American species. A reclassification and nomenclatural changes for the subtribe that are consistent with the phylogeny are provided. Three genera are removed from Loxandrina: Aulacopodus Britton, 1940 moved to Pterostichini Bonelli, 1810; Cosmodiscus Sloane, 1907 and Tiferonia Darlington, 1962 moved to Abacetina. Based on the phylogenetic relationships and nomenclatural priority only four genera are recognized in Loxandrina: Cerabilia Laporte, 1867, Zeodera Laporte, 1867, Pediomorphus Chaudoir, 1878, and Oxycrepis Reiche, 1843. All other previously recognized genera are treated as subgenera. The classification change created eight secondary homonyms that are resolved by the proposal of the following: Oxycrepis gebi, replacement name for O. balli (Straneo, 1993); O. amatona, replacement name for O. matoana (Straneo, 1993); O. xiproma, replacement name for O. proxima (Straneo, 1993); O. rasutulis, replacement name for O. suturalis (Straneo, 1993); O. laevinota, replacement name for O. laevicollis (Bates, 1871); O. arvulap, replacement name for O. parvula (Straneo, 1951); O. noaffine, replacement name for O. affinis (Straneo, 1991); O. alutona, replacement name for O. notula (Tschitschérine, 1901). An overview of the morphological characteristics and diagnostic features of Loxandrina taxa is provided. A key and habitus images are provided for identification of genera and subgenera. The possible historical biogeography of the group is discussed in light of their phylogenetic relationships and past geological events.

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Research Article Wed, 2 Sep 2020 14:23:40 +0300
Anchored between heaven and earth – a new flightless brown lacewing from Peru (Neuroptera, Hemerobiidae) https://dez.pensoft.net/article/56008/ Deutsche Entomologische Zeitschrift 67(2): 141-149

DOI: 10.3897/dez.67.56008

Authors: Ulrike Aspöck, Horst Aspöck, Axel Gruppe

Abstract: Male and female of Nusalala peruana sp. nov., a flightless hemerobiid from the Andes mountain range of northern Peru, at a height of almost 4000 m, are described, figured and documented as the first record of a brachypterous, flightless species of Nusalala Navás, 1913, from this country. The other two congeneric, brachypterous species are from high altitudes in Colombia and Costa Rica and have been described in the male sex only – the females remain unknown. The coriaceous domed forewings are shared by all three brachypterous Nusalala species. The ribbon-like hindwings of the male of N. peruana sp. nov. are unique, since those of the other brachypterous males are scale-like, as are the hindwings of the female of N. peruana sp. nov. Distribution and evolutionary backgrounds of brachyptery and flightlessness in Neuropterida are discussed.

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Research Article Fri, 14 Aug 2020 15:49:15 +0300
Recent additions to the list of German sawflies (Hymenoptera, Symphyta) https://dez.pensoft.net/article/54002/ Deutsche Entomologische Zeitschrift 67(2): 127-139

DOI: 10.3897/dez.67.54002

Authors: Andrew Liston, Marko Prous

Abstract: Two tenthredinid sawfly species are newly recorded in Germany: Pristiphora krausi (Lacourt, 2006) and P. melagonia sp. nov. The latter is also recorded from Greece and Spain. These species belong to the Pristiphora depressa group, as confirmed by genetic data. Additional data are presented for seven other tenthredinid species which have only rarely been recorded in Germany and for which previously-published data are incomplete. Identification keys to the West Palaearctic species of Eurhadinoceraea and the Pristiphora depressa group are included. Other species of Symphyta, first recorded in Germany since the publication of the most recent checklist, are listed in a table, with references to literature which describes identification characters.

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Research Article Tue, 21 Jul 2020 14:30:54 +0300
The Nepalese species of the genus Enicospilus Stephens, 1835 (Hymenoptera, Ichneumonidae, Ophioninae): a preliminary revision and identification key to species https://dez.pensoft.net/article/51332/ Deutsche Entomologische Zeitschrift 67(1): 69-126

DOI: 10.3897/dez.67.51332

Authors: So Shimizu

Abstract: A total of 10 species of Enicospilus (Hymenoptera, Ichneumonidae, Ophioninae) have previously been reported from Nepal. Six new species are described here (E. alleni Shimizu sp. nov., E. kakanicus Shimizu sp. nov., E. nepalensis Shimizu sp. nov., E. nikami Shimizu sp. nov., E. phulchokiensis Shimizu sp. nov., and E. tangi Shimizu sp. nov.), and 10 are newly recorded (E. ashbyi Ashmead, 1904, E. bifasciatus (Uchida, 1928), E. capensis (Thunberg, 1824), E. flavocephalus (Kirby, 1900), E. formosensis (Uchida, 1928), E. grammospilus (Enderlein, 1921), E. pudibundae (Uchida, 1928), E. purifenestratus (Enderlein, 1921), E. yonezawanus (Uchida, 1928), and E. zebrus Gauld & Mitchell, 1981) from Nepal. A preliminary identification key to the Nepalese species of Enicospilus is provided. The elevational pattern of Nepalese Enicospilus is briefly discussed. Enicospilus purifenestratus is also recorded for the first time from Brunei.

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Research Article Mon, 11 May 2020 09:27:28 +0300
A comparative description of the mesosomal musculature in Sphecidae and Ampulicidae (Hymenoptera, Apoidea) using 3D techniques https://dez.pensoft.net/article/49493/ Deutsche Entomologische Zeitschrift 67(1): 51-67

DOI: 10.3897/dez.67.49493

Authors: Maraike Willsch, Frank Friedrich, Daniel Baum, Ivo Jurisch, Michael Ohl

Abstract: Conflicting hypotheses about the relationships among the major lineages of aculeate Hymenoptera clearly show the necessity of detailed comparative morphological studies. Using micro-computed tomography and 3D reconstructions, the skeletal musculature of the meso- and metathorax and the first and second abdominal segment in Apoidea are described. Females of Sceliphron destillatorium, Sphex (Fernaldina) lucae (both Sphecidae), and Ampulex compressa (Ampulicidae) were examined. The morphological terminology provided by the Hymenoptera Anatomy Ontology is used. Up to 42 muscles were found. The three species differ in certain numerical and structural aspects. Ampulicidae differs significantly from Sphecidae in the metathorax and the anterior abdomen. The metapleural apodeme and paracoxal ridge are weakly developed in Ampulicidae, which affect some muscular structures. Furthermore, the muscles that insert on the coxae and trochanters are broader and longer in Ampulicidae. A conspicuous characteristic of Sphecidae is the absence of the metaphragma. Overall, we identified four hitherto unrecognized muscles. Our work suggests additional investigations on structures discussed in this paper.

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Research Article Mon, 11 May 2020 09:25:08 +0300
A review of Himalcercyon stat. nov., with description of a new species from the Chinese Himalaya and an updated key to Asian genera of Megasternini (Coleoptera, Hydrophilidae) https://dez.pensoft.net/article/50078/ Deutsche Entomologische Zeitschrift 67(1): 35-49

DOI: 10.3897/dez.67.50078

Authors: Fenglong Jia, Zulong Liang, Martin Fikáček

Abstract: Himalcercyon Hebauer, 2002 stat. nov. is elevated to genus rank based on the unique form of its mesoventral elevation. The genus is reviewed, redescribed, and illustrated in detail. Two species are recognized: Himalcercyon mirus (Hebauer, 2002) comb. nov. from Nepal and H. franzi sp. nov. from Chinese Himalaya (Xizang Autonomous Region). Both species are illustrated and diagnosed. An updated key to the Asian genera of the tribe Megasternini (Coleoptera, Hydrophilidae, Sphaeridiinae) is provided, along with the SEM micrographs of ventral morphology of these genera. New replacement name Oreosternum nom. nov. is proposed for Oreocyon Hebauer, 2002 which is preoccupied by Oreocyon Marsh, 1872 (Mammalia, Oxyenidae) and Oreocyon Krumbiegel, 1949 (Mammalia, Canidae).

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Research Article Mon, 11 May 2020 08:37:32 +0300
Molecular taxonomy of Tomares hairstreaks (Lepidoptera, Lycaenidae, Theclinae) https://dez.pensoft.net/article/50252/ Deutsche Entomologische Zeitschrift 67(1): 19-33

DOI: 10.3897/dez.67.50252

Authors: Vazrick Nazari, Wolfgang ten Hagen

Abstract: Tomares hairstreaks comprise about 10 species distributed from Europe and North Africa to Central Asia. The taxonomy of the genus is hampered by the absence of diagnostic characters by which specimens can be unambiguously assigned to species. Our investigation of morphology and DNA barcode variations within and between Tomares species shows that while well-defined species (T. ballus, T. mauritanicus, T. callimachus, T. desinens and T. fedtschenkoi) diverge, poorly characterized taxa (T. nogelii, T. nesimachus, T. dobrogensis, T. romanovi and T. telemachus) show very little to no differentiation in mtDNA. We reinstate Tomares callimachus spp. hafis (Kollar, 1849) as a valid subspecies (stat. rev.) and propose taxa telemachus Zhdanko, 2000 and uighurica Koçak, Seven & Kemal, 2000 as synonyms of T. romanovi and T. nogelii nogelii respectively (syn. nov.). We relegate Polyommatus epiphania Boisduval, 1848, recently revived as a valid subspecies of T. callimachus, back to synonymy under the latter, and reconsider the status of T. nogelii dobrogensis (Caradja, 1895) in the light of new molecular data. We use a nuclear gene (EF-1α) in addition to COI barcodes to reconstruct the phylogeny of the group.

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Research Article Tue, 5 May 2020 08:37:12 +0300
Biology of two European Tenthredo species (Hymenoptera, Tenthredinidae) feeding on Gentiana https://dez.pensoft.net/article/49741/ Deutsche Entomologische Zeitschrift 67(1): 13-18

DOI: 10.3897/dez.67.49741

Authors: Andrew Liston, Ewald Altenhofer, Romana Netzberger, Marko Prous

Abstract: Very few sawflies using Gentianaceae as larval host plants have been recorded. We identified larvae collected in Austria on Gentiana asclepiadea L. as Tenthredo atra Linnaeus, 1758 and T. propinqua Klug, 1817. If its current taxonomic circumscription as a single species is accepted, T. atra is a highly polyphagous species, whereas T. propinqua may be more specialised: Gentiana asclepiadea is its first recorded host. We sequenced plant DNA from the head of one T. propinqua larva, which confirmed that it had been feeding on this plant. This is the first recorded use of G. asclepiadea by sawfly larvae. Larvae are illustrated, and identification characters are described.

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Research Article Fri, 21 Feb 2020 10:41:07 +0200
The Tamarix feeding Leafhopper genus Opsius Fieber, 1866 (Hemiptera, Cicadellidae, Deltocephalinae, Opsiini) in the Kingdom of Saudi Arabia, with description of a new species https://dez.pensoft.net/article/46662/ Deutsche Entomologische Zeitschrift 67(1): 1-12

DOI: 10.3897/dez.67.46662

Authors: Saad A. El-Sonbati, Michael R. Wilson, Hathal M. Al Dhafer

Abstract: The leafhopper genus Opsius Fieber, 1866 is revised for the Kingdom of Saudi Arabia. Seven species are treated, including three that previously were reported by Dlabola (1979), O. pallasi (Lethierry, 1874), O. tigripes (Lethierry, 1876), and O. versicolor (Distant, 1908). Opsius heydeni (Lethierry, 1876), O. richteri Dlabola 1960, and O. scutellaris (Lethierry, 1874) are reported for the first time from the Kingdom. A new species, O. wilsoni El-Sonbati, sp. nov. is described from the southwestern region of the Kingdom of Saudi Arabia. A key to the species of Opsius of the Kingdom is also provided.

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Research Article Tue, 14 Jan 2020 09:32:39 +0200
Species composition and diagnoses of leaf- and fruit-scarring beetles (Coleoptera, Chrysomelidae) infesting bananas and plantains (Zingiberales, Musaceae) in the Indian subcontinent https://dez.pensoft.net/article/47447/ Deutsche Entomologische Zeitschrift 66(2): 179-202

DOI: 10.3897/dez.66.47447

Authors: Kaniyarikkal D. Prathapan, Janakiraman Poorani, S. Amritha Kumari, C. Anuradha, Balakrishnan Padmanaban, Ramalingam Thanigairaj

Abstract: Leaf- and fruit-feeding chrysomelids (Coleoptera) on bananas and plantains (Musaceae, Zingiberales) cause major losses to banana growers in the northern and northeastern regions of India, Bangladesh, and other parts of Southeast Asia. The species composition of these beetles has not been studied so far in India and wrong names or wrong name combinations in the literature have caused confusion. Most particularly, the Central and South American apecies of Colaspis hypochlora Lefèvre (Chrysomelidae, Eumolpinae) has been erroneously reported as occurring in India and Bangladesh, and this name has been used for the Indian species. Based on extensive surveys for leaf- and fruit-feeding chrysomelids in the northern and northeastern regions of India from 2015 to 2019, three species of banana-feeding chrysomelids, namely, Basilepta subcostata (Jacoby) (Eumolpinae), Bhamoina varipes (Jacoby), and a new species, Sphaeroderma cruenta sp. nov. (Galerucinae, Alticini), are documented. Of these, the latter two are recorded as pests of banana in India for the first time. An illustrated diagnostic account of these three species is given to facilitate their identification by economic entomologists. COI sequences of populations of B. subcostata from Assam and Uttar Pradesh showed 98–100% homology, indicating that these populations are conspecific and that COI sequences can be used for rapid species determination. Brief notes on the biology and available management options for these pests are also given.

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Research Article Fri, 13 Dec 2019 09:23:29 +0200
Revision of Tropopterus Solier: A disjunct South American component of the Australo-Pacific Moriomorphini (Coleoptera, Carabidae) https://dez.pensoft.net/article/38022/ Deutsche Entomologische Zeitschrift 66(2): 147-177

DOI: 10.3897/dez.66.38022

Authors: James K. Liebherr

Abstract: Tropopterus Solier, 1849, precinctive to southern South America, is taxonomically revised. Six new species are described: T. peckorum sp. nov., T. robustus sp. nov., T. canaliculus sp. nov., T. trisinuatus sp. nov., T. minimucro sp. nov., and T. fieldianus sp. nov. Merizodus catapileanus Jeannel, 1962, is synonymized with T. montagnei Solier, 1849. Lectotypes are designated for T. montagnei, T. giraudyi Solier, T. duponchelii Solier, and T. nitidus Solier (= T. duponchelii). Tropopterus peruvianus Straneo is noted as a nomen dubium, with its identity and taxonomic placement to be substantiated via neotype designation. Phylogenetic relationships among Tropopterus spp. are hypothesized based on 37 morphological characters, the distributions of which are analyzed under the parsimony criterion, with the cladogram root established between Tropopterus and its adelphotaxon from New South Wales, Australia. Speciation in the group has occurred predominantly at a limited geographical scale relative to the overall generic distribution, with three pairs of sister species sympatric. However phylogenetic divergence between taxa in the more northern, sclerophyllous forest characterized by Nothofagus obliqua (Brisseau de Mirbel) and those occupying the Valdivian and North Patagonian Rain Forest dominated by N. dombeyi (Brisseau de Mirbel) is observed in two instances of phylogenetic history. Using specific collecting locality records, it is shown that Tropopterus beetles have been collected syntopically and synchronically with species of Glypholoma Jeannel (Coleoptera, Staphylinidae), Anaballetus Newton, Švec & Fikáček (Coleoptera, Leiodidae), Andotypus Spangler (Coleoptera, Hydrophilidae), and Novonothrus Balogh (Acari, Oribatida). These concordant ecological occurrences document a cohesive Nothofagus forest leaf-litter community. These genera plus other Valdivian Rain Forest invertebrate taxa all exhibit an Austral disjunct biogeographical pattern that corroborates trans-Antarctic vicariance between the Nothofagus forests of southern South America and Australia. Male genitalic antisymmetry is shown to be a synapomorphy of Tropopterus, though the female reproductive tract retains the plesiomorphic orientation observed in all other moriomorphine taxa.

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Research Article Tue, 5 Nov 2019 16:22:11 +0200
A revolutionary protocol to describe understudied hyperdiverse taxa and overcome the taxonomic impediment https://dez.pensoft.net/article/34683/ Deutsche Entomologische Zeitschrift 66(2): 119-145

DOI: 10.3897/dez.66.34683

Authors: Sarah Meierotto, Michael J. Sharkey, Daniel H. Janzen, Winnie Hallwachs, Paul D. N. Hebert, Eric G. Chapman, M. Alex Smith

Abstract: Here we elucidate and justify a DNA barcode approach to insect species description that can be applied to name tens of thousands of species of Ichneumonoidea and many other species-rich taxa. Each description consists of a lateral habitus image of the specimen, a COI barcode diagnosis, and the holotype specimen information required by the International Code of Zoological Nomenclature. We believe this approach, or a slight modification of it, will be useful for many other underdescribed hyperdiverse taxa, especially in the tropics. Due to the extreme species-richness of the Ichneumonoidea, the very low percentage of described species, and the lack of detailed biological information for most described species, the standard taxonomic approach is inefficient and overwhelmingly time consuming. A DNA barcode-based approach to initial description will provide a solid foundation of species hypotheses from which more comprehensive descriptions can be developed as other data, time, and budgets permit. Here we elucidate this view and detailed methodology that can generally be applied to species-rich underdescribed taxa. A real example is given by describing species in two genera, Hemichoma and Zelomorpha, reared from the Área de Conservación Guanacaste in northwestern Costa Rica. The generic type species Zelomorpha arizonensis is given a DNA barcode diagnosis and the following new species are described: Zelomorpha angelsolisi, Zelomorpha bobandersoni, Zelomorpha danjohnsoni, Zelomorpha donwindsori, Zelomorpha effugia, Zelomorpha johnchemsaki, Zelomorpha kellyanneae, Zelomorpha larrykirkendalli, Zelomorpha mariyavladmirovnae, Zelomorpha mikeiviei, Zelomorpha myricagaleae, Zelomorpha noahjaneae, Zelomorpha paulgoldsteini, Zelomorpha terryerwini, Zelomorpha willsflowersi, Hemichoma donwhiteheadi, Hemichoma frankhovorei, and Hemichoma johnkingsolveri.

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Research Article Thu, 25 Jul 2019 09:32:13 +0300
First known larva of omicrine genus Psalitrus d’Orchymont (Coleoptera, Hydrophilidae) https://dez.pensoft.net/article/34300/ Deutsche Entomologische Zeitschrift 66(1): 107-118

DOI: 10.3897/dez.66.34300

Authors: Yûsuke N. Minoshima

Abstract: The larval morphology of the water scavenger beetle Psalitrus yamatensis Hoshina & Satô, 2005 is described based on a specimen collected from Fukuoka Prefecture, Kyushu, Japan. This is the first description of the larval morphology of the hydrophilid genus Psalitrus d’Orchymont, 1919, as well as the first description of larval chaetotaxy of the tribe Omicrini. Species-level identification of the larva was performed using DNA barcoding of a molecular marker: a 658 bp fragment of the mitochondrial cytochrome oxidase I. A description including chaetotaxy of head capsule and head appendages, diagnosis, and illustrations of the larva is provided. Psalitrus larvae can be distinguished from other known larvae of the tribe Omicrini by the morphology of the head and legs. The larva shares characters with other known larvae of Omicrini; potential plesiomorphies are shared with Cylominae and aquatic hydrophilids; some characters are also shared with larvae of Megasternini and Sphaeridiini.

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Research Article Tue, 25 Jun 2019 09:15:36 +0300
Contribution to the knowledge of male and female eremochaetid flies in the late Cretaceous amber of Burma (Diptera, Brachycera, Eremochaetidae) https://dez.pensoft.net/article/33914/ Deutsche Entomologische Zeitschrift 66(1): 75-83

DOI: 10.3897/dez.66.33914

Authors: Qingqing Zhang, Junfeng Zhang

Abstract: A new and a previously known species of the genus Zhenia Q. Zhang, 2016 (Eremochaetidae) are illustrated and described based on two males and a female in amber: Zhenia burmensis sp. nov. and Z. xiai Q. Zhang, 2016. The male Z. xiai is the first male of this species recorded. The relationships of Archisargoidea (including Eremochaetidae, Zhenia) are reassessed based on male genitalia. The superfamily is more likely related to the Stratiomyomorpha than to the Muscomorpha (including Nemestrinoidea). The components and structures of the ovipositor are re-illustrated. The results of our comparative study demonstrate that the ovipositor of Zhenia is similar in shape and detail to that of Rhagoletis pomonella (Walsh, 1867) (Tephritidae). This study concludes that the ovipositor of Zhenia is most likely formed from abdominal eighth and ninth segments instead of the cerci, as a previous study found.

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Research Article Fri, 14 Jun 2019 10:05:35 +0300
On Bulgarian sawflies, including a new species of Empria (Hymenoptera, Symphyta) https://dez.pensoft.net/article/34309/ Deutsche Entomologische Zeitschrift 66(1): 85-105

DOI: 10.3897/dez.66.34309

Authors: Andrew Liston, Marko Prous, Jan Macek

Abstract: Thirty-nine species of sawfly (Symphyta) are recorded for the first time in Bulgaria. Most of these were collected during early spring of 2018, in the south-east of the country (Burgas and Varna Provinces). Empria aridicola Macek & Prous, sp. nov. is described as new to science from specimens collected in several central, east and south European countries. Lectotypes are designated for Poecilosoma parvula Konow, 1892, Empria pravei Dovnar-Zapolskij, 1925 and E. pseudoklugi Dovnar-Zapolskij, 1929. Empria pravei and Sciapteryx byzantina Benson, 1968 are at present only known in Europe from the coastal zone of the Black Sea. The new Bulgarian records of Hoplocampa cantoti Chevin, 1986 and Neomessa steusloffi (Konow, 1892) represent large extensions in their recorded ranges, previously comprising respectively only northern France, and north-eastern Germany. Possible host plant associations are noted for several species, based on observations of adults.

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Research Article Fri, 14 Jun 2019 09:24:26 +0300
The Rubens morph of Formica exsecta Nylander, 1846 and its separation from Formica fennica Seifert, 2000 (Hymenoptera, Formicidae) https://dez.pensoft.net/article/34868/ Deutsche Entomologische Zeitschrift 66(1): 55-61

DOI: 10.3897/dez.66.34868

Authors: Bernhard Seifert

Abstract: A study of numeric morphology-based alpha-taxonomy (NUMOBAT) considering the species Formica exsecta Nylander, 1846 and F. fennica Seifert, 2000 was performed in 166 nest samples with 485 worker individuals originating from 117 localities of the Palaearctic west of 59°E. The presence of intraspecific pilosity dimorphism is shown for F. exsecta. The setae-reduced phenotype, termed the Rubens morph, shows a frequency of about 25%, and the more abundant setae-rich phenotype, termed the Normal morph, one of 75%. The frequency of nests containing workers of both phenotypes is 15.5% in 58 samples from Denmark, Sweden, and Finland. Applying the DIMORPH test of Seifert (2016) on this territory, it is demonstrated that the association of Rubens and Normal phenotypes within the same nest cannot be interpreted as parabiosis of independent species (p=0.017) or as temporary (p=0.0004) and permanent (p=0.0001) socially parasitic association, whereas genetically mediated intraspecific dimorphism is most likely (p=0.659, all p data according to Fisher’s exact test). The Rubens morph of F. exsecta is phenotypically most similar to F. fennica but is safely separable by four different forms of exploratory data analyses using nest centroids (NC) as input data: NC-Ward, NC-part.hclust, NC-part.kmeans, and NC-NMDS-k-means. Data on zoogeography and the narrow climate niche indicate that F. fennica is unlikely to occur in Norway.

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Research Article Wed, 29 May 2019 16:11:23 +0300
About Notiophilus Duméril, 1806 (Coleoptera, Carabidae): Species delineation and phylogeny using DNA barcodes https://dez.pensoft.net/article/34711/ Deutsche Entomologische Zeitschrift 66(1): 63-73

DOI: 10.3897/dez.66.34711

Authors: Michael J. Raupach, Karsten Hannig, Jérome Morinière, Lars Hendrich

Abstract: The genus Notiophilus Duméril, 1806 is a distinctive taxon of small, diurnal and morphologically similar beetles exhibiting large eyes and widened second elytral intervals. In this study we analysed the effectiveness of DNA barcodes to discriminate 67 specimens that represent 8 species of Notiophilus from Central Europe. Interspecific K2P distances below 2.2% were found for N. biguttatus (Fabricius, 1779) and N. quadripunctatus Dejean, 1826, whereas intraspecific distances with values > 2.2% were revealed for N. rufipes Curtis, 1829. An additional phylogenetic analysis of all available species revealed a close relationship of N. directus Casey, 1920, N. semistriatus Say, 1823, N. simulator Fall, 1906 and N. sylvaticus Dejean, 1831, possibly indicating a radiation of these species in North America. Low support values of most other nodes, however, do not allow additional phylogenetic conclusions.

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Research Article Wed, 29 May 2019 15:25:55 +0300
Type catalogue of the thick-headed flies (Diptera, Conopidae) in the collection of the Museum für Naturkunde Berlin https://dez.pensoft.net/article/33814/ Deutsche Entomologische Zeitschrift 66(1): 41-53

DOI: 10.3897/dez.66.33814

Authors: Jens-Hermann Stuke, Joachim Ziegler

Abstract: Type material of thick-headed flies (Diptera, Conopidae) in the collection of the Museum für Naturkunde Berlin (ZMHB) is documented. The entire collection holds primary type material (i.e. holotypes, lectotypes, syntypes) of 73 species. Five Conops species with previously unknown subgenus belong to the subgenus Asiconops: C. frontosus Kröber, 1916; C. indicus Kröber, 1916; C. maculiventris Kröber, 1916; C. nigrofasciatus Kröber, 1916; and C. punctifrons Kröber 1916. Two new synonyms are introduced: Conops vaginalis Rondani, 1865 syn. nov. of Conops truncatus Loew, 1847 and Siniconops fuscatus Qiao & Chao, 1998 syn. nov. of Physocephala sepulchralis Brunetti, 1912.

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Research Article Wed, 15 May 2019 12:19:33 +0300
Redescriptions of thirteen species of chewing lice in the Brueelia-complex (Phthiraptera, Ischnocera, Philopteridae), with one new synonymy and a neotype designation for Nirmus lais Giebel, 1874 https://dez.pensoft.net/article/32423/ Deutsche Entomologische Zeitschrift 66(1): 17-39

DOI: 10.3897/dez.66.32423

Authors: Daniel R. Gustafsson, Lucie Oslejskova, Tomas Najer, Oldrich Sychra, Fasheng Zou

Abstract: Thirteen species of chewing lice in the Brueelia-complex are redescribed and illustrated. They are: Brueelia blagovescenskyi Balát, 1955, ex Emberiza schoeniclus (Linnaeus, 1758); B. breueri Balát, 1955, ex Chloris chloris (Linnaeus, 1758); B. conocephala (Blagoveshchensky, 1940) ex Sitta europaea (Linnaeus, 1758); B. ferianci Balát, 1955, ex Anthus trivialis (Linnaeus, 1758); B. glizi Balát, 1955, ex Fringilla montifringilla Linnaeus, 1758; B. kluzi Balát, 1955, ex Fringilla coelebs Linnaeus, 1758; B. kratochvili Balát, 1958, ex Motacilla flava Linnaeus, 1758; B. matvejevi Balát, 1981, ex Turdus viscivorus Linnaeus, 1758; B. pelikani Balát, 1958, ex Emberiza melanocephala Scopoli, 1769; B. rosickyi Balát, 1955, ex Sylvia nisoria (Bechstein, 1792); B. vaneki Balát, 1981, ex Acrocephalus schoenobaenus (Linnaeus, 1758); Guimaraesiella haftorni (Balát, 1958) ex Turdus iliacus Linnaeus, 1758; G. lais (Giebel, 1874) ex Luscinia megarhynchos (Brehm, 1831). Redescriptions are made from type material where available. Holotypes are identified in Balát’s material when possible, and lectotypes are designated for B. blagovescenskyi, B. breueri, B. glizi, B. ferianci, B. kluzi, B. kratochvili, B. pelikani, and B. rosickyi; a neotype of Nirmus lais Giebel, 1874 is designated. Brueelia weberi Balát, 1982, is placed as a synonym of Brueelia conocephala (Blagoveshchensky, 1940).

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Research Article Mon, 4 Feb 2019 09:58:30 +0200
Playing hard to get: two new species of subterranean Trechini beetles (Coleoptera, Carabidae, Trechinae) from the Dinaric Karst https://dez.pensoft.net/article/31754/ Deutsche Entomologische Zeitschrift 66(1): 1-15

DOI: 10.3897/dez.66.31754

Authors: Roman Lohaj, Teo Delić

Abstract: Almost 200 years of continuous and systematic research in subterranean habitats of the Dinaric Karst and adjoining areas have resulted in the discovery of more than 400 specialized subterranean beetles. Among these, a special place belongs to the morphologically well distinguished and elusive, so called aphaenopsoid trechine beetles, which are characterized by a prolonged head, pronotum and appendages, and widened, ovoid-shaped elytra. Two new species of aphaenopsoid trechines – Derossiella lukici sp. n. from two deep pits on Mt Biokovo, Croatia, and Adriaphaenops petrimaris sp. n. from Pištet 4 Cave, Kameno more, Montenegro – are described, illustrated, and compared with closely related congeners. Identification keys for both genera and an annotated catalogue for all Adriaphaenops species, as well as data on the distribution and the ecology of these remarkable species, are provided and discussed.

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Research Article Mon, 21 Jan 2019 09:39:18 +0200
Revision of Zosteragathis Sharkey of Thailand (Hymenoptera, Braconidae, Agathidinae, Agathidini) https://dez.pensoft.net/article/25772/ Deutsche Entomologische Zeitschrift 65(2): 225-253

DOI: 10.3897/dez.65.25772

Authors: Michael J. Sharkey, Eric G. Chapman

Abstract: Based on cladistic analyses recently conducted by Sharkey and Chapman, the genus Zosteragathis Sharkey is revised. Twenty-two species are reported from Thailand, three previously described species, Z. samensis, Z. contrasta and Z. nuichuaensis, and 19 new species, i.e., Z. chaiyaphumensis, Z. eukos, Z. hinensis, Z. hongensis, Z. inthanonensis, Z. krachanensis, Z. lampangensis, Z. lampooensis, Z. luangensis, Z. ngamensis, Z. perknos, Z. petchaburiensis, Z. phahompokensis, Z. phuphanensis, Z. sakaeratensis, Z. sakonensis, Z. samensis, Z. surinensis, Z. taemensis, Z. tonensis. Members of Zosteragathis are known from the Australian, Ethiopian, Oceania, Oriental, and eastern Palaearctic regions.

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Research Article Tue, 14 Aug 2018 15:18:09 +0300
Taxonomic review of Australian Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) with special emphasis on the M. lophoides (Chaudoir) species complex https://dez.pensoft.net/article/27424/ Deutsche Entomologische Zeitschrift 65(2): 177-224

DOI: 10.3897/dez.65.27424

Authors: James K. Liebherr

Abstract: The Australian fauna of Mecyclothorax Sharp (Coleoptera: Carabidae: Moriomorphini) is reviewed, with special focus on species assigned to the monophyletic subgenus Eucyclothorax Liebherr: M. isolatus, sp. n. from Western Australia, M. moorei Baehr, M. punctatus (Sloane), M. curtus (Sloane), M. blackburni (Sloane); M. eyrensis (Blackburn); M. peryphoides (Blackburn); M. darlingtoni, sp. n. from Queensland; M. jameswalkeri, sp. n. from Western Australia; M. lophoides (Chaudoir); and M. cordicollis (Sloane). The last six species listed above–the M. lophoides species complex–have been the source of long-term confusion for taxonomists, with male genitalic characters providing trouble-free species circumscription. One new subspecies, M. lewisensis estriatus, subsp. n. from Queensland is added to the seven previously described taxa of the monophyletic subgenus Qecyclothorax Liebherr. The balance of the fauna consists of four species in the subgenus Mecyclothorax: 1 and 2, the sister-species pair M. lateralis (Castelnau) and M. minutus (Castelnau); 3, M. ambiguus (Erichson); and 4, M. punctipennis (MacLeay). Mecyclothorax fortis (Blackburn), syn. n., is newly synonymized with M. minutus. Mecyclothorax ovalis Sloane is recombined as Neonomius ovalis (Sloane), comb. n., and a neotype is designated to replace the destroyed holotype. Phylogenetic relationships for the Australian Mecyclothorax are proposed based on information from 68 terminal taxa and 139 morphological characters. The biogeographic history of Australian Mecyclothorax is deduced based the sister-group relationship between Mecyclothorax and the Amblytelus-related genera, with both groups hypothesized to have originated during the late Eocene. Diversification within Mecyclothorax has occurred since then in montane rainforests of tropical Queensland, temperate forest biomes of the southwest and southeast, and in grasslands and riparian habitats adjacent and inland from those forests. Several species presently occupy interior desert regions, though no sister species mutually occupy such climatically harsh habitats. The M. lophoides species complex exhibits profound male genitalic diversification within the context of conserved external anatomy. This disparity is investigated with regard to the functional interaction of the male internal sac flagellum and female spermathecal duct. Though limited association of flagellar and spermathecal duct configurations can be documented, several factors complicate proposing a general evolutionary mechanism for the observed data. These include: 1, the occurrence of derived, elongate spermathecal ducts in three species, two of which exhibit very long male flagella, whereas males of the third exhibit a very short flagellum; and 2, a highly derived and exaggerated male flagellar configuration shared across a sister-species pair even though the two species can be robustly diagnosed using external anatomical characters, other significant genitalic differences involving male parameral setation, and biogeographic allopatry associated with differential occupation of desert versus forest biomes.

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Research Article Mon, 13 Aug 2018 13:47:59 +0300
The tropical African genus Morgenia (Orthoptera, Tettigoniidae, Phaneropterinae) with emphasis on the spur at the mid tibia https://dez.pensoft.net/article/26693/ Deutsche Entomologische Zeitschrift 65(2): 161-175

DOI: 10.3897/dez.65.26693

Authors: Bruno Massa, Klaus-Gerhard Heller, Elżbieta Warchałowska-Śliwa, Nicolas Moulin

Abstract: The authors revised the genus Morgenia Karsch, 1890 which now consists of eight species, of which three are here newly described (Morgenia plurimaculata Massa & Moulin, sp. n., M. angustipinnata Massa, sp. n., and M. lehmannorum Heller & Massa, sp. n.). Six of the eight species occur in the Tri National Sangha (TNS) comprising Dzanga-Sangha Special Reserve and Dzanga Ndoki National Park (Central African Republic), whose high biodiversity has been recently highlighted. In particular the genus is characterised by the presence of a more or less long spur at the inner mid tibia, different in each species; in M. modulata, it moved lower down into a new position at about ¼ of tibia, which has a hollow underneath where the rest of the spur remains hidden. This is a unique known case in Phaneropterinae. Morphological characters distinguishing males of different species are presented. Bioacoustics of the new species M. lehmannorum are described. The patterns of the chromosome evolution in M. lehmannorum differ from other investigated African Phaneropterinae in terms of chromosome number and morphology, reduced ancestral chromosome number (2n = 25) implying a more derived condition.

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Research Article Thu, 2 Aug 2018 09:14:20 +0300
Revision of the Quedius fauna of Middle Asia (Coleoptera, Staphylinidae, Staphylininae) https://dez.pensoft.net/article/27033/ Deutsche Entomologische Zeitschrift 65(2): 117-159

DOI: 10.3897/dez.65.27033

Authors: Maria Salnitska, Alexey Solodovnikov

Abstract: Twenty eight species of the genus Quedius from Middle Asia comprising Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan and Uzbekistan, are revised. Quedius altaicus Korge, 1962, Q. capitalis Eppelsheim, 1892, Q. fusicornis Luze, 1904, Q. solskyi Luze, 1904 and Q. cohaesus Eppelsheim, 1888 are redescribed. The following new synonymies are established: Q. solskyi Luze, 1904 = Q. asiaticus Bernhauer, 1918, syn. n.; Q. cohaesus Eppelsheim, 1888 = Q. turkmenicus Coiffait, 1969, syn. n., = Q. afghanicus Coiffait, 1977, syn. n.; Q. hauseri Bernhauer, 1918 = Q. peneckei Bernhauer, 1918, syn. n., = Q. ouzbekiscus Coiffait, 1969, syn. n.; Q. imitator Luze, 1904 = Q. tschinganensis Coiffait, 1969, syn. n.; Q. novus Eppelsheim, 1892 = Q. dzambulensis Coiffait, 1967, syn. n., Q. pseudonigriceps Reitter, 1909 = Q. kirklarensis Korge, 1971, syn. n. Lectotypes are designated for Q. asiaticus Bernhauer, 1918, Q. fusicornis Luze, 1904, Q. hauseri Bernhauer, 1918, Q. imitator Luze, 1904, Q. novus Eppelsheim, 1892 and Q. solskyi Luze, 1904. For all revised species, taxonomy, distribution and bionomics are summarized. Quedius fuliginosus (Gravenhorst, 1802), Q. sundukovi Smetana, 2003 and Q. pseudonigriceps Reitter, 1909 are recorded for Middle Asia for the first time. One species from the Q. coloratus-group, found to be new to science is not described due to shortage of material. Another possibly new species is tentatively identified as Q. fulvicollis Stephens, 1833 until the taxonomy of that widespread species is revised. An identification key to all species is provided.

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Research Article Tue, 31 Jul 2018 10:05:58 +0300
Review of the genus Tricerophora Janse, 1958 (Lepidoptera, Gelechiidae) with description of six new species https://dez.pensoft.net/article/25747/ Deutsche Entomologische Zeitschrift 65(1): 81-98

DOI: 10.3897/dez.65.25747

Authors: Oleksiy V. Bidzilya, Wolfram Mey

Abstract: An improved diagnosis of the genus Tricerophora is provided, and its position within Gelechiidae is briefly discussed. A new generic synonym is established: Leucophylla Janse, 1960, syn. n. of Tricerophora Janse, 1958. Six new species are described: T. pundamilia sp. n. (RSA), T. rukinga sp. n. (Kenya), T. nigrinervis sp. n. (RSA, Namibia), T. brumale sp. n. (Namibia), T. acutivalva sp. n. (Iran), T. minimorum sp. n. (Namibia). The following new combinations are proposed: Tricerophora nigribasis (Janse, 1960), comb. n., Tricerophora objecta (Meyrick, 1921), comb. n. A key to the species is given based on external characters and the genitalia of both sexes. Adults and genitalia of all species are illustrated.

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Research Article Tue, 12 Jun 2018 14:18:41 +0300
Review of the flower-inhabiting water scavenger beetle genus Cycreon (Coleoptera, Hydrophilidae), with descriptions of new species and comments on its biology https://dez.pensoft.net/article/26261/ Deutsche Entomologische Zeitschrift 65(1): 99-115

DOI: 10.3897/dez.65.26261

Authors: Emmanuel Arriaga-Varela, Sin Yeng Wong, Alexander Kirejtshuk, Martin Fikácek

Abstract: The hydrophilid genus Cycreon Orchymont, 1919, previously known from two historical specimens only, is reviewed based on the numerous material collected recently from the inflorescences of various Araceae species in the Malay Peninsula and Borneo. Four species are recognized in the genus: C. sculpturatus Orchymont, 1919 from Sumatra, C. armandi Shatrovskiy, 2017 from Singapore, C. adolescens sp. n. from peninsular Malaysia, and C. floricola sp. n. with two subspecies, the nominotypical one from Peninsular Malaysia, and C. floricola borneanus subsp. n. from Borneo. All species are very similar, differing only by the pronotal punctation, shape of the clypeus and the mentum, and the form of the median lobe of the aedeagus. Specimens of C. floricola sp. n. and C. adolescens sp. n. were collected from inflorescences of various genera of the family Araceae. The field observations and analysis of mid gut contents indicates that they feed on organic material on internal organs of the inflorescences, including the pollen of the host plant. They were also observed to carry a large amount of pollen and are likely pollinators of their host species of Araceae.

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Research Article Tue, 12 Jun 2018 10:13:01 +0300
Vansoniella chirindensis gen. n., sp. n. – an unusual taxon with translucent wings from Zimbabwe (Lepidoptera, Limacodidae) https://dez.pensoft.net/article/23538/ Deutsche Entomologische Zeitschrift 65(1): 75-80

DOI: 10.3897/dez.65.23538

Authors: Wolfram Mey

Abstract: The genus Vansoniella gen. n. is established to accommodate the species V. chirindensis sp. n., collected in Zimbabwe by Van Son in 1937. The new species differs externally from other African taxa by translucent fore- and hindwings in the male sex. The wing venation is highly derived and the male genitalia are also structurally different from other genera. The genus occupies an isolated position within the family.

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Research Article Fri, 16 Mar 2018 09:22:00 +0200
Mouthpart dimorphism in male and female wasps of Vespula vulgaris and Vespula germanica (Vespidae, Hymenoptera) https://dez.pensoft.net/article/23593/ Deutsche Entomologische Zeitschrift 65(1): 65-74

DOI: 10.3897/dez.65.23593

Authors: Bianca Baranek, Kenneth Kuba, Julia Bauder, Harald Krenn

Abstract: Social wasps perform a variety of tasks with their mouthparts. Female workers use them to feed on carbohydrate-rich fluids, to build nests by collecting wood fibers and forming paper, to hunt and manipulate insect prey for feeding larvae as well as for brood care. Since male wasps neither feed on insects nor participate in nest building, sex-specific differences in mouthpart morphology are expected. Despite these different applications, general mouthpart morphology of male and female wasps from the genus Vespula was similar. However, males possessed significantly shorter mandibles with fewer teeth than females. Furthermore, the adductor muscles of the mandibles were distinctly smaller in males than in females. Male wasps showed a higher number of sensilla on the mandibles and the labial palpi. Mouthpart dimorphism and functional morphology of fluid uptake are discussed.

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Research Article Tue, 13 Mar 2018 10:36:49 +0200
Cladistic classification of Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) and taxonomic revision of the New Caledonian subgenus Phacothorax Jeannel https://dez.pensoft.net/article/21000/ Deutsche Entomologische Zeitschrift 65(1): 1-63

DOI: 10.3897/dez.65.21000

Authors: James K. Liebherr

Abstract: The 15 species of Mecyclothorax Sharp precinctive to New Caledonia are revised and shown by cladistic analysis to comprise a monophyletic lineage, here treated as subgenus Phacothorax Jeannel. The New Caledonian species of subgenus Phacothorax include Mecyclothorax fleutiauxi (Jeannel), M. najtae Deuve, and 13 newly described species: M. jeanneli sp. n., M. laterobustus sp. n., M. laterorectus sp. n., M. laterosinuatus sp. n., M. laterovatulus sp. n., M. manautei sp. n., M. megalovatulus sp. n., M. octavius sp. n., M. paniensis sp. n., M. picdupinsensis sp. n., M. plurisetosus sp. n., and two jointly authored species; M. kanak Moore & Liebherr sp. n., and M. mouensis Moore & Liebherr sp. n.. subgenus Phacothorax is one of five subgenera recognized within genus Mecyclothorax based on cladistic analysis of 65 exemplar taxa utilizing information from 137 morphological characters. The four other monophyletic subgenera include the precinctive Australian Eucyclothorax subgen. n. (type species Mecyclothorax blackburni [Sloane]), the precinctive Queensland Qecyclothorax subgen. n. (type species Mecyclothorax storeyi Moore), the precinctive New Zealand Meonochilus Liebherr & Marris status n., and the geographically widespread and very diverse nominate subgenus, distributed from St. Paul and Amsterdam Islands, eastward across Australia and New Guinea, and in the Sundas, Timor Leste, Lord Howe and Norfolk Islands, New Zealand, and the Society and Hawaiian Islands. The biogeographic history of Mecyclothorax can be derived from the parsimony cladogram time-calibrated by times of origin of particular geographic areas inhabited by resident representative species. Based on sister-group status of subgenus Phacothorax and subgenus Mecyclothorax, and occupation of Lord Howe Island–an island originating no earlier than 6 Ma–by the earliest divergent lineage within subgenus Mecyclothorax, the ancestor of present-day Phacothorax spp. is hypothesized to have colonized New Caledonia 6 Ma, subsequent both to Cretaceous Gondwanan vicariance as well as any Oligocene submergence. Area relationships among the New Caledonian Phacothorax point to earliest diversification incorporating the northern massifs, and most recent diversification on the ultramafic volcanic substrates in the south of Grand Terre. Flight wing loss has played an important role in shaping the various island faunas, both in their morphology as well as their diversity. The retention of flight capability in only a few of the many hundred Mecyclothorax spp. is presented in light of how populations evolve from macropterous colonizing propagules to vestigially winged specialists. Interspecific differences in genitalic structures for the sister-species pair M. fleutiauxi + M. jeanneli are shown to involve functional complementarity of male and female structures. Extensive geographic variation of male genitalia is demonstrated for several New Caledonian Mecyclothorax spp. This variation deviates from the geographically uniform male genitalia exhibited by species in the hyperdiverse Mecyclothorax radiation of Haleakalā volcano, Maui, suggesting that extensive sympatry occurring among species in that diverse species swarm selects for stability within this mate recognition system. Conversely, lower levels of sympatry characterizing the depauperate New Caledonian radiation permit the presence of more extensive male genitalic variation, this variation not selected against due to the lower likelihood of interspecific mating mistakes.

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Research Article Thu, 18 Jan 2018 10:42:46 +0200
160 years of D.E.Z. – what is the recipe for thy long life? https://dez.pensoft.net/article/22742/ Deutsche Entomologische Zeitschrift 64(2): 161-161

DOI: 10.3897/dez.64.22742

Authors: Dominique Zimmermann

Abstract: This year we look back at 160 years of entomological research published in the DEZ. Believe it or not, our journal is the third oldest of all still existing entomological periodicals worldwide! A concatenation of favourable circumstances? At first glance, the first decades were rather tough ones, involving personal controversies, splitting of the society behind the journal and the journal itself, and later reunion (Wessel 2007). However, at the second glance, this period seems to have been an excellent one at the same time, as the young and dedicated visionary, Gustav Kraatz, the first editor of the DEZ, guided the journal throughout these troublesome waters for the first 50 years. What makes him visionary? Already 160 years ago, he promoted high standards in taxonomical publications such as the description of both sexes when erecting new genera, the publication of comprehensive revisions instead of single species descriptions and the exploration of new diagnostic characters (Wessel 2007) – not much to add 160 years later! More than this, under his editorship the DEZ was at the forefront of the development and establishing of internationally recognized nomenclatorial rules in entomology, regulating foremost issues of priority (Wessel 2007). Still today, nomenclatorial issues constitute a hot topic in entomological publishing.Finally, Gustav Kraatz was driven by the urge to combine collections and libraries of all German entomologists, so that scientists could have free access (Wessel 2007). In 1886, he founded an Entomological National Museum that still exists and is nowadays known as DEI – Deutsches Entomologisches Institut. It would have certainly pleased Gustav Kraatz that since the transfer of the DEZ from Wiley to Pensoft in 2014 all articles are published under an open access policy, likewise facilitating the access to knowledge.The incredible number of 22.613 species descriptions published in the last 160 years in the DEZ (Stelbrink and Wessel 2008; numbers updated) are a substantial contribution to our knowledge of the insects on this planet. The vision of Gustav Kraatz and the commitment of many following editors and authors have made the success of the DEZ possible. Together let’s do our best to continue this line to the future!

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Editorial Thu, 7 Dec 2017 12:17:52 +0200
Revised taxonomic check list of the Eurasiatic species of the subtribe Poliina (Noctuidae, Noctuinae, Hadenini) https://dez.pensoft.net/article/21455/ Deutsche Entomologische Zeitschrift 64(2): 133-160

DOI: 10.3897/dez.64.21455

Authors: Zoltán Varga, Gábor Ronkay, László Ronkay

Abstract: The revised checklist of the subtribe Poliina Hampson, 1902 is presented; one new genus, Multisignagen. n., three new subgenera (Atropolia, Leuconephropolia and Protopoliasubgen. n.) and a new species (Polia (Atropolia) posterodilutasp. n.) are described. The taxonomic position of the recently described subgenus Metallopolia is discussed. The subtribe Pachetrina Beck, 1996 is synonymised with Poliina; two genera (Kollariana Hacker, 1996 and Spiramater McCabe, 1980) are transferred to the subtribe Mamestrina Hampson, 1902. A number of lectotype designations and new combinations are given; the newly designated lectotypes and the genitalia of the disputed taxa are illustrated.

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Review Article Thu, 30 Nov 2017 09:01:47 +0200
Anthropogenic dispersal of a snakefly (Insecta, Neuropterida) – a singular phenomenon or a model case in Raphidioptera? https://dez.pensoft.net/article/19859/ Deutsche Entomologische Zeitschrift 64(2): 123-131

DOI: 10.3897/dez.64.19859

Authors: Horst Aspöck, Ulrike Aspöck, Axel Gruppe, Marcia Sittenthaler, Elisabeth Haring

Abstract: The Mediterranean snakefly Raphidia mediterranea H. Aspöck, U. Aspöck & Rausch, 1977 – known from many parts of the Balkan Peninsula, several Aegean islands, southern parts of Italy, northwest of Anatolia and a few localities in Eastern Europe, yet not recorded in Central Europe – was surprisingly found with an astoundingly high population density on bushes in the yard of an old farmhouse at a comparatively high altitude (800 m) in Upper Austria, north of the Danube River, in 2013. This spectacular phenomenon was again observed in the following years (2014, 2015, 2016, 2017), and in 2016 the suspicion that the larvae develop in the straw of the thatched roof of the farmhouse could be confirmed by findings of larvae, pupae, and exuviae.It is most likely, that the occurrence of this Raphidia species in Austria is to be traced back to a human-caused introduction at some point in time. It remains, however, unknown when and specifically how this event might have occurred. Morphologically no substantial differences were found between specimens from Greece, Italy and Upper Austria. In addition, the genetic uniformity (using 3 genes: cox1, cox3, and 28S) of the populations was verified. This supports an earlier hypothesis that the occurrence of the species, as well as that of R. mediterranea in Italy, Anatolia and perhaps elsewhere, may be related to importation of goods involving wood or soil. A molecular genetic analysis of several Raphidia species confirmed the present morphology-based concept of their systematic position. The means of dispersal of Raphidioptera are largely unknown. We do not know of any other similar cases of anthropogenic dispersal of a snakefly, but it cannot be excluded that human activities may have played a greater role in the dispersal of Raphidioptera than previously assumed. Phylogenomic studies would therefore be promising to solve some of these questions.

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Research Article Mon, 2 Oct 2017 11:44:26 +0300
New findings of Flagellisargus J Zhang, 2012 (Diptera, Brachycera, Archisargidae), with discussion of the placements of some controversial taxa https://dez.pensoft.net/article/13550/ Deutsche Entomologische Zeitschrift 64(2): 111-122

DOI: 10.3897/dez.64.13550

Authors: Junfeng Zhang

Abstract: A new species of a new subgenus and a similar known species referred to the genus Flagellisargus J Zhang, 2012 are described and illustrated based on a male and a female impression fossils of these flies: Flagellisargus (Changbingisargus) parvussubgen. et sp. n. and Flagellisargus (Flagellisargus) cf. sinicus J Zhang, 2012. The latter taxon is the first record of a female Flagellisargus. Recently taken out of Archisargoidea, this study concludes that Flagellisargus should be an archisargid genus based on the known (male) and new (female) impression fossils. The placement of Daohugosargus J Zhang, 2012b is reassessed. It demonstrates close similarities in body structure and wing venation to archisargid flies, and can be retained as an archisargid genus. Archirhagio mostovskii J Zhang, 2015 is separated from Archirhagio zhangi K Zhang et al., 2009. Helempis Ren, 1998 could be, as a separate genus, placed in Archisarginae, Archisargidae.

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Research Article Thu, 3 Aug 2017 09:48:09 +0300
The Nevrorthidae, mistaken at all times: phylogeny and review of present knowledge (Holometabola, Neuropterida, Neuroptera) https://dez.pensoft.net/article/13028/ Deutsche Entomologische Zeitschrift 64(2): 77-110

DOI: 10.3897/dez.64.13028

Authors: Ulrike Aspöck, Horst Aspöck, Xingyue Liu

Abstract: This monographic review of the Nevrorthidae Nakahara, 1915, covers all 19 validly described, extant species worldwide that belong to one of the smallest families of the order Neuroptera. The family embraces four genera: Nevrorthus Costa, 1863 (with five species occurring in the Mediterranean region), Austroneurorthus Nakahara, 1958 (with two species restricted to eastern Australia), Nipponeurorthus Nakahara, 1958 (with 11 species from eastern Asia: Japanese islands, mainland China, Taiwan), and Sinoneurorthus Liu, H. Aspöck & U. Aspöck, 2012 (with one species recorded from mainland China). A comprehensive taxonomical treatment of all extant taxa is presented, including the scant available biological data. Distribution maps for all species are provided. A phylogenetic analysis based on morphological data from both extant and extinct taxa was performed. Austroneurorthus, together with Nevrorthus and some Eocene Baltic amber genera, form a monophylum. The disjunct distribution of modern nevrorthid genera demonstrates the relictual nature of the family and points to a historical biogeography that could have led to the formation of the present distribution pattern. Future discovery of fossil material might substantiate these claims. Sonnet for a Vulnerable Creature Is the Climate still fine? Still clean, the Riverine? Ruined rivulets run dry Fossils – tho’ living – may Die.

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Research Article Thu, 3 Aug 2017 09:09:46 +0300
Pseudochrysis Semenov, 1891 is the valid genus name for a group of cuckoo wasps frequently referred to as Pseudospinolia Linsenmaier, 1951 (Hymenoptera, Chrysididae) https://dez.pensoft.net/article/13005/ Deutsche Entomologische Zeitschrift 64(1): 69-75

DOI: 10.3897/dez.64.13005

Authors: Paolo Rosa, Maurizio Pavesi, Villu Soon, Oliver Niehuis

Abstract: The cuckoo wasp genus Pseudochrysis Semenov, 1891 is currently treated by several authors as a junior subjective synonym of Euchroeus Latreille, 1809, due to a type species designation by O. W. Richards in 1935. In the original description of the genus Pseudochrysis, Semenov (1891) distinguished two subordinated taxa within the genus Pseudochrysis: the subgenus Pseudochrysis and the subgenus Spintharis (sensu Dahlbom 1854). Semenov included three species in the subgenus Spintharis, but failed to mention any species included in the nominal subgenus. He was the first author, however, who listed in a subsequent publication (Semenov 1892) eleven species to be included in the nominal subgenus. According to the International Code of Zoological Nomenclature (ICZN 1999, Art. 67.2.2), these eleven species are deemed to have been listed in the original description. One of these, Chrysura humboldti Dahlbom, 1845, was explicitly designated by Semenov (1892) as type species of Pseudochrysis. We therefore consider the designation of Pseudochrysis (Spintharis) virgo Semenov, 1891 as type species of Pseudochrysis by Richards (1935) as invalid. The currently widely used genus name Pseudospinolia Linsenmaier, 1951 (type species Chrysis uniformis Dahlbom, 1854) is consequently to be regarded as a junior subjective synonym of Pseudochrysis, given the current circumscription of the genus Pseudospinolia (including both Pseudospinolia humboldti and Pseudospinolia uniformis).

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Short Communication Wed, 31 May 2017 10:07:27 +0300
On the enigmatic Sinonemestrius Hong & Wang, 1990, with description of a new species based on a complete fossil fly (Diptera, Brachycera, Tabanomorpha, Heterostomidae) https://dez.pensoft.net/article/11724/ Deutsche Entomologische Zeitschrift 64(1): 61-67

DOI: 10.3897/dez.64.11724

Authors: Junfeng Zhang

Abstract: A new species of Sinonemestrius Hong & Wang, 1990 is described and illustrated based on a complete compression fossil of the fly. Currently placed in Rhagionemestriidae or Xylophagidae, the present study concludes that the morphology of the new find indicates that Sinonemestrius is a heterostomid genus within Tabanomorpha. The placement of Ahirmoneura neimengguensis K-y Zhang et al., 2008 is reassessed. It demonstrates close similarities in body structure and wing venation to those of Sinonemestrius, and can be provisionally transferred from Nemestrinidae to Heterostomidae: Sinonemestriinae.

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Research Article Fri, 7 Apr 2017 11:31:15 +0300
Taxonomy and phylogeny of the genera Gymnocnemia Schneider, 1845, and Megistopus Rambur, 1842, with remarks on the systematization of the tribe Nemoleontini (Neuroptera, Myrmeleontidae) https://dez.pensoft.net/article/11704/ Deutsche Entomologische Zeitschrift 64(1): 43-60

DOI: 10.3897/dez.64.11704

Authors: Davide Badano, Horst Aspöck, Ulrike Aspöck

Abstract: The delineation of antlion genera has often been based on morphological characters not tested in a phylogenetic context, thus seriously impairing the study of systematics of the family Myrmeleontidae. Nebulous generic limits also impede the taxonomy and study of the affinities of closely related species. As a case study, the generic placement of Megistopus mirabilis Hölzel, 1980, was based on a single leg character. To test the position of this species, the reciprocal relationships of the members of the genera Gymnocnemia Schneider, 1845, and Megistopus Rambur, 1842 were investigated, using a morphology-based phylogenetic analysis. This approach demonstrated that M. mirabilis should actually be assigned to the genus Gymnocnemia, as G. mirabilis comb. n. This analysis also supports the subdivision of the tribe Nemoleontini in two subclades based on morphology of male and female genitalia. A new characterisation of these genera is provided, as well as a redescription of the very rare G. mirabilis and the poorly investigated Megistopus lucasi (Navás, 1912). An updated identification key to the members of the genera Gymnocnemia and Megistopus is presented.

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Research Article Wed, 8 Mar 2017 09:51:28 +0200
Morphology of the larvae of three Central European Strophosoma Billberg, 1820 (Coleoptera, Curculionidae, Entiminae) species https://dez.pensoft.net/article/11446/ Deutsche Entomologische Zeitschrift 64(1): 27-42

DOI: 10.3897/dez.64.11446

Authors: Rafal Gosik, Peter Sprick, Katarzyna Czerewko

Abstract: The larvae of Strophosoma (Strophosoma) capitatum (DeGeer, 1775), S. (Strophosoma) melanogrammum (Forster, 1771) and S. (Neliocarus) sus Stephens, 1831, are illustrated and re-described or described for the first time. The first larval instar, and the mature or an older larval instar, are illustrated, and a general description of the Strophosoma larva is given. The biological data obtained from breeding and field-collecting are compared and discussed in relation to the known life-cycle data.

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Review Article Tue, 21 Feb 2017 10:39:23 +0200
Revisions of the Afrotropical genera of Argidae and species of Pampsilota Konow, 1899 (Hymenoptera, Tenthredinoidea) https://dez.pensoft.net/article/10800/ Deutsche Entomologische Zeitschrift 64(1): 1-25

DOI: 10.3897/dez.64.10800

Authors: Andrew D. Liston, Georg Goergen, Frank Koch

Abstract: The Afrotropical fauna contains five genera of Argidae. These are keyed. New subjective synonyms, followed by the valid name in brackets, are Calarge Enslin, 1911 [Arge Schrank, 1802], Calarge africana Enslin, 1911 [Arge congrua Konow, 1907], Clyparge Pasteels, 1963 [Scobina Lepeletier & Serville, 1828], Clyparge terminalis Pasteels, 1963 [Scobina poecila (Klug, 1834)], and Sterictophora [sic] afra Pasteels, 1963 [Sphacophilus afer comb. n., species inquirenda near S. monjarasi Smith & Morales-Reyes, 2015]. The type material of both C. terminalis and S. afra was probably collected in the New World, but labelled with the wrong locality “Kamerun”. An introduction of both species to Africa, not followed by long-term establishment, seems less likely. The removal of these taxa from the faunal list of the region is recommended. The nine known Afrotropical species of Pampsilota are revised, and an illustrated dichotomous identification key presented, with distribution maps for all species. Four species are here described as new to science: P. dahomeyanus Goergen, Koch & Liston, sp. n., P. nigeriae Liston & Koch, sp. n., P. tsavoensis Liston & Koch, sp. n., and P. zebra Liston & Koch, sp. n. Lectotypes are designated for Pampsilota afer Konow, 1899, and Cipdele africana Mocsáry, 1909. The immature stages and host plant of only one species are known: P. dahomeyanus on Lannea nigritana (Anacardiaceae). Its larval morphology strongly resembles that of European and North American species of Arge. We provisionally retain Pampsilota as a valid genus, although it could justifiably be treated as comprising merely a species group, or groups, within Arge.

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Research Article Thu, 19 Jan 2017 12:53:54 +0200
Hidden species within the genus Ocys Stephens: the widespread species O. harpaloides (Audinet-Serville) and O. tachysoides (Antoine) (Coleoptera, Carabidae, Bembidiini) https://dez.pensoft.net/article/10748/ Deutsche Entomologische Zeitschrift 63(2): 287-301

DOI: 10.3897/dez.63.10748

Authors: David R. Maddison, Roy Anderson

Abstract: Beetles previously considered to be Ocys harpaloides (Audinet-Serville) from northern Africa, Spain, France, the United Kingdom, France, and Belgium belong to two species. These species can be distinguished using DNA sequences of 28S rDNA, 18S rDNA, COI, CAD, and topoisomerase I. A key, diagnoses, and images are provided to enable identification of specimens based upon characteristics of male and female genitalia, as well as microsculpture and other external structures. Through examination of the holotype of Bembidium harpaloides v. tachysoides Antoine, as well as designation of lectotypes of Bembidion harpaloides Audinet-Serville and Ocys melanocephalus Stephens, and designation of a neotype for Tachis rufescens Guérin-Ménéville, the valid names of the two species were determined to be Ocys harpaloides and Ocys tachysoides (Antoine).

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Research Article Wed, 30 Nov 2016 10:17:09 +0200
Spotted males, uniform females and the lowest chromosome number in Tettigoniids recorded: Review of the genus Gonatoxia Karsch (Orthoptera, Phaneropterinae) https://dez.pensoft.net/article/10799/ Deutsche Entomologische Zeitschrift 63(2): 271-286

DOI: 10.3897/dez.63.10799

Authors: Claudia Hemp, Klaus-Gerhard Heller, Elzbieta Warchalowska-Sliwa, Andreas Hemp

Abstract: The genus Gonatoxia Karsch, which was synonymized with Dapanera Karsch by Massa (2015), is re-established. Data on habitat, biology, ecology, the acoustics and on chromosomes are provided as well as a key to the species. The male of G. immaculata Karsch and the female of G. maculata Karsch are described. G. furcata sp. n. from the Udzungwa Mountains of Tanzania and G. helleri sp. n. inhabiting coastal and lowland wet forest are newly described. All species have calling songs consisting of very short, resonant syllables, produced in species specific intervals, and with peak carrier frequencies between 13 and 24 kHz. In respect to chromosome numbers Gonatoxia species showed extreme differences (from 29 to 7) suggesting rapid evolutionary changes. G. helleri sp. n. so far is the tettigoniid species with the lowest number of chromosomes at present. Gonatoxia species may be used as bioindicators, their presence suggesting valuable habitats that are vanishing rapidly in East Africa.

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Research Article Fri, 25 Nov 2016 12:02:06 +0200
Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia https://dez.pensoft.net/article/10241/ Deutsche Entomologische Zeitschrift 63(2): 211-270

DOI: 10.3897/dez.63.10241

Authors: James Liebherr

Abstract: The precinctive New Caledonian genus Cyphocoleus Chaudoir is revised with 22 species recognized, 12 newly described: C. lissus sp. n., C. prolixus sp. n., C. parovicollis sp. n., C. burwelli sp. n., C. angustatus sp. n., C. monteithi sp. n., C. fasciatus sp. n., C. lescheni Liebherr & Will, sp. n., C. cordatus sp. n., C. bourailensis sp. n., C. subulatus sp. n., and C. iledespinsensis sp. n. Atongolium Park & Will is found to be a junior synonym of Cyphocoleus, with its two species recombined as C. mirabilis comb. n. and C. moorei comb. n. Results of a survey of Harpalinae Bonelli place Cyphocoleus as a member of Odacanthini based on synapomorphies of the eighth abdominal tergite and the female spermathecal assembly. Cyphocoleus shares with five other generic-level taxa – Homethes Newman, Aeolodermus Andrewes, Stenocheila Laporte, Quammenis Erwin and Diplacanthogaster Liebke – a single-segmented maxillary galea that is appressed to the outer margin of the maxillary lacinia. These six generic-level taxa are newly classified as members of subtribe Homethinasubtrib. n. (type genus Homethes). Cladistic analysis including 79 taxa and utilizing 119 morphological characters supports division of Odacanthini into four monophyletic subtribes: 1, Actenonycina (Actenonyx White); 2, Homethina; 3, Pentagonicina (Pentagonica Dana, Parascopodes Darlington, Scopodes Erichson); and 4, Odacanthina (24 genera in this analysis monophyletically defined by Lasiocera Dejean and its adelphotaxon). These subtribes are phylogenetically arranged as: (Actenonycina (Homethina (Pentagonicina + Odacanthina). Area relationships defined within Homethina – (New Caledonia (Australia (South America + Central America))) – support the origin of New Caledonian Cyphocoleus prior to amphiantarctic vicariance between South America and Australia. Consistent with previous molecular dating of 100–105 Ma for the origin of Odacanthini, a general vicariance-based hypothesis proposes that New Zealandian Actenonyx and New Caledonian Cyphocoleus were emplaced on Zealandia prior to the completion of rifting between Zealandia and Australia during Late Cretaceous, and that both fragments of Zealandia remained subaerial throughout the Cenozoic. Alternatively, under a very specific time-constrained biogeographic hypothesis ladened with an added assumption of dispersal, the ancestor of Cyphocoleus could have colonized New Caledonia during a 2–5 Ma period after its proposed subaerial reemergence at 37 Ma. A clade within Cyphocoleus synapomorphously exhibits an environmental patina: a varnish-like coating to the dorsal body surface that is hypothesized to enable crypsis of the adult beetle. Several specializations of elytral setae are also synapomorphies of this clade, suggesting evolutionary association of the patina and the setal specializations.

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Research Article Fri, 18 Nov 2016 10:38:44 +0200