Krell F, Philips T (2010) Formicdubius Philips & Scholtz from South Africa, a junior synonym of Haroldius Boucomont, and a survey of the trichomes in the African species (Coleoptera, Scarabaeidae, Onthophagini). ZooKeys 34: 41-48. doi: 10.3897/zookeys.34.342